122 I\[r. TI. ?,. Fantliara on 



fig. ], (J.) attracted attention as soon as seen, being yellowish 

 brown in colour, rather elongate ovoid in shape, and slightly 

 larger, darker, and more pointed anteriorly than the testes in 

 normal male specimens. The gonads were approximately equal 

 in size and general shape, about 1'9 cm. long, I'l cm. broad, 

 and 1 cm. thick. Especially noteworthy was the presence of 

 a conspicuous yellow e^g (fig. 1, o.) on the ventral surface of 

 the posterior third of the left gonad (hence use of term 

 *' ovotestis '' for this gonad). This egg was '3 cm. in diameter, 

 but those found in females of T. grceca were often larger. 



The Miillerian ducts were well developed and lay ventral 

 and to the outer side of each gonad (fig. 1, m.). Tiiey were 

 light yellow in colour and approximately equal in length, and 

 their external diameter varied from '3 cm. anteriorly to 

 ■6 cm. in the uterine portion. They were attached by a 

 mesentery posteriorly to the epididymis of either side, and 

 anteriorly by the broad ligament to the lungs. Each opened 

 in front by an ostium, apparently ciliated (fig. 1, Fl.) , into 

 the ccelom, considerably in front of the gonads. No eggs, 

 however, were noticed free in the body-cavity, nor in the 

 Miillerian ducts, the lumen of each of the latter being con- 

 tinuous throughout. These oviducts opened into the neck of 

 the cloacal bladder, slightly behind and ventral to the ureters 

 and vasa deferentia, as shown in PI. VI. fig. 1. Tlie 

 foldings of these ducts were chiefly confined to their anterior 

 third. 



Tortoise B. — This specimen exhibited all the external 

 characters of the male, the concavity of its plastron being 

 well marked. The genitalia (PI. VI. fig. 2) were of the 

 normal size, shape, and appearance common to the male 

 tortoise, each testis being 1"6 cm. long, 1*2 cm. broad, and 

 1 cm. thick. But in addition there were rudimentary 

 Miillerian ducts present (fig. 2, »?.), opening anteriorly into 

 the coelom, that of the left side being slightly longer than that 

 of the right, but rather narrower in its posterior sixth ajid 

 at its entrance to the cloaca (PI. VI. fig. 2). The hinder 

 portion of the right oviduct being wider ("3 cm. in external 

 diameter), its posterior fifth was found to contain functionally 

 mature spermatozoa, which, however, were not found in the 

 more constricted hinder part of the left oviduct. Apparently 

 the basal part of the right rudimentary oviduct functioned as 

 a sort of vesicula seminalis {cf. Howes^s observations [6] on a 

 specimen of Lacerta viridis), the spermatozoa having pro- 

 bably migrated into it from the vas deferens. Unfortunately 

 the oviducts of specimen A were not examined for spermatozoa 

 when fresh, for reasons given below\ 



