

into v/hicli ihe splits fimilly open, loavinf^ tlie sorcil 

 canals tliuG open to tlie exterior for a time, to "be clococl 

 ci^ciin later by the ^-rov/in^^ top;ether of the cells at the 

 vontral sui'Iace* On liic outer v/alls of these cavities 

 arise the "soFal cells", in eacJi of which walls appear later 



so as to cut out a tetrtiliedral apical cell. This cuts 

 off a number of SG(!,inents, .vhich according to Paisso^; ^^ive 



rise to the placenta v/ith its vascular bundle, cind to the 

 microsporangia, then ci pcricline appears at the inner end 

 of the apical cell forminr: thus the archesporium of the 

 single nacrosporangiujn arising from it. Goebel ('82) 

 states tliat the soral Ccinal^ of Mars ilia are external in 

 origin and that the sportmgia a.rise from superficial cells. 

 Bftsgen ('90) described the earliest stage of the sporocarp 

 is in one case a "gap" (Lf^cke) in the epidei-mal tissue of 

 the young leaf near its base, and in another place as a 

 swelling or prominence in the Scime region. Tlie sporocea^p 

 grows for a time by a two sided c^pical cell, and he thinks 

 it probable that all of the soral colls are derived from 

 an epidermal cell of the ventral side of the capsule. 

 Tlie placenta macrospcrL-oigia and microsporangia ho states 

 are all foiTned as Russow has shov,-n from these soral cells. 



In the similfir sporocarp of Pilularia, Hofmeister 

 (•62) cind Juranyi ('79) describe the soral cavities as 

 arisinp-, internally, while Croobcl ('o?J, Meunier ('C?) imd 



