9/ 



The viev/s of 0LTI.IAII1TS and YAJIAITOUCHI coincide so fc>,r 

 as regarding the sporoeenous cells ol' the cystocarp as "be- 

 longing to the sporophytic phase of an antithetic alterna- 

 tion of generations. Tlie point of departure lies in the 

 interpretation of the tetraspore-producing plant. Because 

 of its general morphological identity v/ith the gametophyte, 

 OLTTIAIIMS regards this as a part of the gai/ietophyte , vhich 

 has become differentiated for the production of tetraspores. 

 Because of the diploid condition of its nuclei, Yamanouchi 

 regards the tetrasporic plant as a part of the sporophyte, 

 whose resemblance to the gametophyte is stamped on it by 

 "similar environmenta,l conditions". 



For the purposes of the present discussion, I sliall 

 assume from the c^-^tological evidence what it will take cul- 

 tural experiments to prove, nai.iely, tlia-t in those red al- 

 gae in which tetraspores and gametes are regularly formed 

 on separate individuals there is an actual succession of 

 sexual and tetrasporic plants, the reproductive bodies of 

 one kind of plant always producing tlie other kind of plant. 



YAIIAJ'OUCKI ' S suggestion ('06b) that the tetrasporic 

 plant may iiave arisen jihylogenetically b:' tiie postponement 

 of the phenomena of reduction from the formation of carpo- 

 spores to tlie production of asexual spores seems to be re- 



