The Relation of Organic to Mental Evolution. 265 



and alterations of its machinery during the lifetime of the 

 individual. How otherwise could an animal have any 

 power of individual acquisition ? But while heredity pro- 

 vides a power of individual acquisition, it does not provide 

 in advance for particular innovations or modifications. A 

 store of innate power, and definite susceptibilities to 

 pleasure and pain, are inherited ; individual experience 

 utilizes this innate power to mould its instinctive and 

 other activities into new habits under the guidance of 

 pleasure and pain. And this absence of provision in 

 advance for particular modifications is probably what 

 Komanes meant. Apart from this, he seems to have quite 

 clearly indicated a valid criterion of the effective presence 

 of consciousness narnety, the observable, profiting by 

 experience. It is just because a chick, having tasted a 

 nasty grub, refuses to touch it or its like for the future, 

 that we may feel confident he exercises real choice in the 

 matter. Invariable response under similar conditions, 

 as in the case of the spermatozoids in the presence of 

 malic acid, is no evidence of choice ; but an alteration or 

 modification of response in the light of individual 

 experience does afford the criterion we seek. And this 

 criterion involves the exercise of control. Automatism, no 

 matter how complex or how nicely adaptive, may be un- 

 conscious ; but controlled action indicates conscious choice. 

 Whatever the physiological conditions of the process 

 may be, it seems clear that consciousness, as choosing and 

 controlling, stands in a sense outside that upon which it 

 exercises control through its power of choice. And, thus, 

 on passing from the merely organic to the conscious- 

 organic phase of evolution, we have to consider the develop- 

 ment of an imperium in imperio. In close touch and 

 intimate relation with the merely organic and automatic 

 system is a controlling system, the functional activity of 



