THE CIRCULATION OF THE BLOOD 



371 



recording apparatus, e.g., a tambour with a lever or a piston recorder with 

 float and writing point. The space between the organ and vessel is filled 

 with normal saline, air, or oil. Such an apparatus is known as a plethysmo- 

 graph. Fig. 175. Many forms of this apparatus have been devised in 

 accordance with the character of the organ spleen, kidney, etc. to be 

 investigated, though the principle underlying them is essentially the same. 

 In addition to changes in volume due to the heart's action, most organs 

 undergo additional changes in volume from vaso-motor and respiratory 

 causes. 



Indeed the plethysmographic is the most generally employed method 

 of showing the action of vaso-motor nerves in changing the contraction of the 

 arterioles and hence the outflow of blood. Thus when an organ is enclosed 



p IG< 175. A PLETHYSMOGRAPH FOR THE ARM. 



in a plethysmograph and the arterial contraction increased by either a direct 

 or reflex stimulation of the vaso-motor center there will be a rise in the 

 pressure, a diminution in the outflow of blood and a decrease in the 

 volume of the organ under observation; and on the contrary, if the artenole 

 contraction is diminished by a direct or reflex inhibition of the vaso-moto 

 center there will be a fall of pressure, an increased outflow of blood and , 

 increase in the volume of the organ. From this it is learned that the func 

 tional activity of an organ which is attended and conditioned by an increased 

 blood-supply is always associated with an increase in volume. On plethys- 

 mographk records large undulations are frequently observed which are 

 regarded as of respiratory origin. 



THE CAPILLARY CIRCULATION 



In certain regions of the body of many animals it is possible, on account 

 of the delicacy and transparency of the tissues, to observe not only the flow 



