32 



THE STRUCTURE OF FLOWERS. 



cords, one at each, end of the long axis, now part company 

 from the rest, and enter the lateral sepals (Fig. 6 (a) l.s.), 

 the antero-posterior sepals next receiving their cords (a.s. 

 and^.s.). The cylinder tends to close up, and four groups 

 situate at the corners of the oblong cylinder supply cords for 

 the i^etals, p. The two honey-glands next put in an appear- 

 ance, G. They are merely cellular expansions of the floral 

 receptacle, and are entirely devoid of cords, and therefore 

 not rudiments of appendages. The two lateral stamens next 

 receive their cords, l.st., while four other cords are given off 

 from beside the petaline for the taller pairs of stamens, st. 



p.s 



a 



ls\ = ( = i.sl 



t\l.s 



Anatomy of Wallflower. 



Fig. 6 (6) shows how their cords diverge below and spring 

 from the siae of the petaline cords, while extra cords arise 

 between them to form the marginal cords of the carpels (m.c). 

 From this it will be seen that the longer stamens cannot be 

 formed by " chorisis " of a common intermediate cord ; but, 

 like those of all other members of the flower, their cords are 

 sepai-ated from the common fibro-vascular cylinder of the 

 stem. 



The conclusion suggested by this investigation, and by a 

 comparative study of Gapparidea', is that a cruciferous flower 

 is not reducible to an originally quaternary type at all, but 

 to some higher one. In my paper referred to, I suggested a 



