240 THE STRUCTUKE OF FLOWERS. 



before the development of the styles and stigmas, bore 

 diminished whitish anthers, not opening at all, and containing 

 also some shrivelled pollen-grains. Lastly, in a barren sandy 

 locality, many of the stems produced female flowers, with 

 stamens aborted in the same degree as in D. superhus, and 

 not infrequently such female flowers and protandrous her- 

 maphrodite ones are found on the same stem." Wiegman also 

 found the Dianthus had contabescent stamens when growing 

 on a dry and sterile bank. The conditions here mentioned 

 are very like those more than once described as associated 

 with double flowers, in which the stamens have also de- 

 generated but taken the petaloid form. Hence I think we 

 may directly trace the degeneracy of the anthers and pollen 

 to atrophy ; since chemical analyses of pollen prove that the 

 most important constituents required are potash, nitrogen, 

 and phosphorus pentoxide,* pi'obably wanting in the localities 

 mentioned. 



" Centaurea Jacea " Midler describes f " as having its 

 flower-heads of the same stem aUvays of the same form, but 

 different stems of the same locality often present astonishing 

 differences in their flower-heads. 



"In the most common and apparently original form, the 

 floAver-heads consist of florets which are all of the same 

 tubular shape, and all contain both fully developed anthers 

 and stigma, the divergence of the outer florets giving to the 

 whole head a diameter of 20-30 mm. From this original 

 form variation has gone on in two opposite dii-ections, the 

 final effects of this variation being, on the one side, very 

 conspicuous male flower-heads of 50-55 mm. diameter ; and 

 on the other side less conspicuous female flower-heads of 



* From an analysis uf Ash blossoms, by Professor Church, Journal 

 of Botany, 1877, p. 36-4. 



f Nature, vol. xxv., p. 2-41. 



