THE GREAT ANTEATER. 305 



with various modifications, iri nearly all the members of this group. 

 Onjcteropus in its cerebral characters seems to approach Myrmecophaga 

 more nearly than any other form, the sulci and gyri of the brains of 

 the two forms, as well as their general conformation, being very similar ; 

 Manis seems to possess the three typical sulci well developed ; and these 

 are also present in the larger Dasypodidae, though apparently much re- 

 duced in the smaller forms of that group. The Sloths conform to the 

 same general type. But, in the absence of a larger series of brains of 

 this group than is at present available for comparison, satisfactory gene- 

 ralization on this subject is impossible, most of the published figures of 

 Edentate brains being very unsatisfactory in detail, whilst nothing of 

 importance is known as regards the development of the sulci in any 

 member of this group. 



3. Female Generative Organs (fig. 6, p. 306). These have been briefly p. z. S. 1882, 

 described by Pouchet *, as well as by Kapp ; but their accounts will, in P- 2ytJ - 

 some respects, bear supplementing. 



A cloaca, in the true sense of the word, is not present in the Great 

 Anteater. The labia majora, which bound the vertical urine-genital 

 fissure, are very prominent and hirsute. Above them, but separated by 

 a distinct perineal space, slightly hair-clad, is the transverse anal aperture, 

 the mucous membrane lining which is pink, quite different from that of 

 the lower passage and its boundaries, which is grey. Slightly inclosing 

 these two apertures above is a widely-open V-shaped tegumentary fold, 

 with its apex situated superiorly towards the root of the tail. 



There are no labia minora visible ; and no clitoris is present as a free 

 organ, though the corpora cavernosa can be felt as tough bodies lying in 

 the walls of the vulva. 



The length of the uriiio-genital canal is 2*7 inches : about 1 inch from 

 its external orifice may be seen, on each side of the middle line, two or 

 three small pore-like depressions ; a bristle passed through the largest of 

 these enters a short duct, connected with one of a pair of globular com- 

 pact glands about the size of a small cherry, which lie in the walls of the 

 urine-genital canal above, between it and the rectum. They are, no 

 doubt, " vulvo-vaginal " glands, or glands of Bertolini, corresponding to 

 the male Cowper's. The urino-genital canal is lined by smooth, vascular, 

 mucous membrane. 



Communication between this and the next section of these organs is 

 effected by means of two small apertures, each admitting readily enough 

 the passage into the vagina, through the here constricted walls of the 

 common tube, of a probe. From between these apertures is prolonged 

 downwards, for a slight distance along the dorsal wall of the urino-genital 

 canal, a slight ridge of mucous membrane, on each side of which are 



* Mem. p. 194. 



