THE CHINESE WATER-DEER. 315 



though not having the complex structure of the ileo-csecal gland met 

 with in Moschus, Cervus, Camelopardalis, &c. 



The only figure hitherto extant (that given by Prof. Grarrod in his 

 paper already quoted) of the brain of Hydropotes having been taken from 

 a very young specimen, it may be worth while to give figures of the 

 superior and lateral aspects of that removed from this adult specimen, 

 which will be useful for comparison with Garrod's earlier one. as well as 

 with those given by that author and Prof. Flower of the brain in Ela- 

 pliodus, Moschus, and Pudua, and with the series of semidiagrammatic 

 sketches illustrating Dr. Krueg's valuable paper on the cerebral convolu- 

 tions of the Ungulata generally*, whose nomenclature on the subject I 

 have also adopted. 



In its cerebral organization Hydropotes approaches the genus Capreolus 

 more nearly than any other Cervine form known to me, the similarity 

 of the two being obvious on comparison of the figures now exhibited 

 (see p. 314) with those of Leuret and Gratiolet t and of Krueg of the 

 Roe. From Elapliodus and Pudua these two forms differ in the entire 

 disappearance (save very slightly anteriorly) of the calloso-marginal 

 (" splenial ") sulcus from the superior aspect of the hemispheres, owing 

 to the greater " pronation " of their brain generally. 



Sir Victor Brooke has been led, from a consideration of other points , 

 to associate Hydropotes and Capreolus with Alces, as a group per se, with 

 affinities in some points in the direction of the Old- World (Plesio- 

 metacarpal), in others in that of the New- World (Idiometacarpal) forms. 

 It appears to me that the additional evidence in this paper, especially 

 that derived from the resemblance of the generative organs, is strongly 

 in favour of this association, so far, at least, as Hydropotes and Capreolus 

 are concerned. The general similarity in fades of Capreolus to Hydro- 

 potes has often struck me, and has even, I believe, led others into the 

 error of mistaking one for the other ! 



That Hydropotes is in no way intimately related to MoscJius was already 

 amply demonstrated ; and the latter form also differs, as we now know, 

 in the conformation of its glans penis and in the possession of Cowper's 

 glands. 



* Zeitschr. f . \vissenschaftl. Zool. xxxi. pp. 297-344. Cf. also Garrod, Coll. Papery 

 pp. 512-517. 



t Anat. Syst. Nerveux, Atlas, pi. x. { L. c. pi. xxi. 



P. Z. S. 1878, p. 889. 



