MACROSCOPIC AND MICROSCOPIC CHARACTERS OF PLANTS. 



793 



very slightly sickle-shaped; they are curved toward the 

 base and curve upwards toward the apex, but this is not 

 so pronounced as in L. purpurata; and there are green 

 nectaries at the apices, though these are not so large as in 

 C. mossice. (Table J 13.) 



The sepals and petals of L. purpurata are white with 

 pale lilac veins; those of C. mossice, pink-lilac; and those 

 of the hybrid, a paler pink-lilac. 



The petals are shorter and narrower in L. purpurata 

 than in C. mossice, and in the hybrid are very nearly mid- 

 intermediate in length and width but slightly nearer 

 C. mossice. (Table J 13.) 



The labellum is very nearly the same length in the 

 parents and hybrid, but is wider in the hybrid than in 

 either parent. (Table J 13.) 



Table J 13. 



Lengths and widths of sepals: 



L. purpurata: Length of dorsal sepals 10.2 em.; of lateral sepals 



10.2 cm.; width of sepals 2.1 cm. 

 C. mossice: Length of dorsal sepals 10.5 cm.; of lateral sepals 



9 cm.; width of sepals 2.3 cm. 

 L.-C. canhamiana: Length of dorsal sepals 10.5 cm.; of lateral 



sepals 9.8 cm.; width of sepals 2.4 cm. 

 Lengths and widths of the petals: 



L. purpurata: Length 9.5 cm.; width 4.4 cm. 

 C. mossice: Length 10.3 cm.; width 5.7 em. 

 L.-C. canhamiana: Length 10 cm.; width 5.2 cm. 

 Lengths and widths of labellum: 



L. purpurata: Length 8.1 cm.; width 6 cm. 



C. mossice: Length 8.2 cm.; width 6 cm. 



L.-C. canhamiana: Length 8 cm.; width 6.7 cm. 



The anterior part of the labellum in L. purpurata 

 has a wavy margin ; that of C. mossice is very wavy, the 

 indentations being comparatively deep ; and that of the 

 hybrid between the parents but slightly nearer L. pur- 

 purata. The tip of the labellum is pointed in L. purpu- 

 rata, and in a few specimens it is slightly indented, about 

 2 mm. deep ; in C. mossice there is quite a deep cleft, on 

 an average 9 mm. deep; in the hybrid the cleft is not 

 quite so deep, about 3 to 4 mm. The upper surface of the 

 basal half of the labellum in L. purpurata is white with 

 a yellowish tinge, and with reddish-purple veins. In 

 C. mossite the basal part (not quite half of the labellum) 

 at the margin is lilac, like the petal and sepals, and 

 a pale yellow between the reddish brownish-purple veins. 

 The rest of the basal half and part of the apical half is 

 yellow, with brownish-purple veins. In the hybrid the 

 basal part (not quite half of the labellum) at the mar- 

 gin is white, and yellowish-white between the red-violet 

 veins. The part corresponding to the yellow area of 

 C. mossice does not include quite so much of the apical 

 half as in C. mossice, it is of a paler shade of yellow, and 

 it is not so distinct an area as in C. mossice, the white 

 basal area blending gradually into the pale yellow area. 

 The apical half of the labellum in L. purpurata is a rich, 

 velvety crimson-purple fading to pinkish-white at the 

 extreme tip ; that of C. mossia is blotched magenta with 

 a distinct pale lilac margin; that of the hybrid has near 

 the tip a blotched magenta area very similar to that of 

 C. mossice, and posterior to this a darker colored area 

 more like the crimson-purple of L. purpurata, becoming 

 paler towards the margin and more of a lavender. 



The column in L. purpurata is white with a few pur- 

 ple dots and purple margins on the anterior face ; that of 

 C. mossice is entirely white; that of the hybrid is white 

 27 



with a few purple specks on the anterior face and suffused 

 with pale pink lilac on the posterior face. (Table J 14.) 



Table J 14. — Lengths and widths of the column. 

 L. purpurata: Length 3 cm.; width 1 cm. 

 C. mossice: Length 4 cm.; width 0.9 cm. 

 L.-C. canhamiana: Length 3 cm.; width 0.95 cm. 



The pollinia in L. purpurata are 8 small masses, 2 by 

 1.3 mm. ; in C. mossue, 4 much larger masses with tails, 

 3 by 2 mm.; and in the hybrid, 4 large masses with tails 

 3 by 2 mm. and 2 small masses 1 by 0.5 mm. 



COMPAHISONS OF THE MICROSCOPIC CHARACTERS. 



Roots. 



The roots of L.-C. canhamiana were in poor condi- 

 tion, on which account studies were not made of this part 

 of the plant. 



PSEUDOBULBS. 



Pseudobulbs of the same age were sectioned at the 

 middle of the thickened part, the third internode (Plate 

 30, figs. 178, 179, and 180). The epidermis in L. pur- 

 purata and the hybrid is composed of rather thick-walled 

 cells that are deeper than wide; those of C. mossice are 

 practically as deep as wide. The depth of the epidermis 

 cells of L. purpurata is much greater than that of C. 

 mossice, while in the hybrid it is only slightly less than in 

 L. purpurata. The width is much less in the hybrid than 

 in either parent. (Table J 15.) 



Table J 15. — Depth and width of cells of epidermis of pseudobulb. 



L. purpurata: Depth 42.5m; width 33.5m- 

 C. mossice: Depth 30.9m; width 31.3m- 

 L.-C. canhamiana: Depth 41m; width 20.2m. 



The outer face of the epidermal cells is provided 

 with a very thick cuticle which is of the same depth 

 (28.8^) in both parents, but much deeper in the hybrid 

 (39.6/*) than in the parents. Beneath the epidermis in 

 L. purpurata are 2 layers of thick-walled, radially elon- 

 gated cells. In C. mossice, the first layer consists of thick- 

 walled, non-elongated cells, and the second layer of cells 

 not quite as thick-walled. In the hybrid there are two 

 layers of cells not quite as thick-walled, nor as elongated, 

 as in L. purpurata. In the hybrid the length of the 

 cells of the first layer is intermediate between the parents 

 and slightly nearer that of C. mossice; they are narrower 

 than in either parent, and in thickness of cell wall inter- 

 mediate but nearer L. purpurata. (Table J 16.) 



Table J 16. — Depth, width, and thickness of walls of first layer 

 of cells beneath the epidermis. 



L. purpurata: Depth 108. 4m; width 38.9m; thickness of walls 10.8 



to 14.4m- 

 C. mossice: Depth 40.7m; width 36m; thickness of walls 3.6m. 

 L.-C. canhamiana: Depth 70.6m; width 28m; thickness of walls 7.2 



to 10.8m. 



Within the thickened hypodermal tissue are the 

 bundles which are embedded in a tissue of large, thin- 

 walled cells. Some of the latter contain needle crystals ; 

 others, starch grains; and others, mucilaginous matter. 



Leaf. 



Sections of the upper epidermis of the leaf were made 

 at the apex, middle, and base of the leaf. The epidermal 

 cells are rectangular, pentagonal, or hexagonal, always 



