CHESTNUT BARK DISEASE. 42! 



here at present, the handwriting is that of Schwaegrichen, and 

 the specimen is an authentic Schweinitzian one. It is probably 

 one of a set sent to Hooker, though there is nothing on the 

 label to indicate that this was the case. There is a pencil refer- 

 ence in another handwriting (apparently Berkeley's) to 'Fr EL 

 2 P- 73- Versatiles.' Some ascospores have been found in this 

 specimen from which the accompanying drawing has been made. 

 They measure 5-7.5x2-3 /A (average size about 7x2 /). The 

 spores are usually one-septate. The septa are indistinct unless 

 stained." 



From Shear's and Wakefield's measurements of the spores, 

 one can readily see that the specimen in the Kew herbarium 

 labeled Sphaeria radicalis is not the species we are considering 

 here under that name, but really the next species, Endothia 

 gyrosa. In a previous publication (9) we stated our belief that 

 Schweinitz's S. radicalis and 5\ gyrosa represented either the 

 two distinct species of Endothia that we now find in the south- 

 ern United States or else the conidial and the asco-stage of 

 only one of them, most likely S. gyrosa. This Kew specimen 

 points to the latter of these two conclusions. It has also been 

 the opinion of certain European botanists that these two species 

 of Schweinitz were merely synonyms, and identical with the 

 form found in Europe, which we call Endothia gyrosa. 



Ellis (N. Am. Pyren. p. 552) in his description included both 

 of these species (his spore measurements relating to one and 

 his drawings to the other), though most of the specimens he 

 referred to are those with linear spores. Farlow (20) was 

 the first to really point out the two as distinct species, and 

 because of this we (9) previously referred to the linear-spored 

 form as Endothia radicalis (Schw.) Farl., though Farlow never 

 definitely used this combination for the fungus. While at 

 present it seems somewhat doubtful if Schweinitz's Sphaeria 

 radicalis really relates to this fungus, we shall retain this com- 

 bination, hoping for further light on the subject through future 

 investigation. On the other hand, there is little if any doubt 

 that Schweinitz's Peziza cinnabarina does relate to its conidial 

 stage, since it is identical, and has Liquidambar for a host, 

 a host upon which E. gyrosa has not yet been reported. The 

 nomenclature already used for this fungus by different writers 

 is as follows : 



