128 



THE BIOLOGY OF A PLANT. 



c. 



the upper part of the leaf, as might be expected from theit 

 functions in connection witli the action of light (see page 147). 



The epidermis^ or shin of the leaf, consists of translucent, 

 greatly flattened cells having peculiar wavy outlines and rela- 

 tively thick walls (Figs. 58-61). Upon 

 the veins they become elongated, and 

 their walls are considerably thickened, 

 especially upon the midrib (Fig. 58, 

 They generally contain large, distinct 

 nuclei, and often considerable proto- 

 j^lasm. The wavy epidermal cells, 

 particularly in young plants, contain 

 some chlorophyll and starch, though 

 1-c* in this respect the fern is somewhat 

 exceptional. 



In the rhizome the epidermis forms 

 a continuous layer over the whole sur- 



(After Sachs.)— Epi- facc. In the leaf, however, this is not 

 dermal cells of ptcris flabei- ^]^g ^^^^ ^l^g epidermis ou the lower 



tola, showing the development ^ ^ ^ ^ 



'• of stomata. A, very young side being perforated by holes leading 

 Z?::-rL.!lima.t,ric^ ^^to the interior and known as months 



mother-cell; sx, sudsidiary or stOlJiata (singular, StomO) (Fig. 61). 

 cell ; g.c, guard-cell ; St, stoma, ^m ii i x -xxi n 



Ihese holes do not pass into the ceils, 

 but are gaps or breaks between certain cells of the epidermis, 

 and open directly into the intercellular spaces, of wliich they are, 

 in fact, the ends. That portion of the intercellular labyrinth 

 which directly underlies the stoma is sometimes called the respira- 

 tory cavity. Each stoma is l)ounded, as in most plants, by two 

 curving guard-cells^ which are generally nucleated, and, unlike 

 epidermal cells generally, contain abundant chlorophyll-bodies 

 and starch. 



The guard-cells are capable of clianging their form accord- 

 ing to the amount of light, the hygroscopic state of the atmos- 

 phere, and other circumstances, and thus open or close the hole 

 or stoma between them. This action is of great importance in 

 the physiology of the plant (transpiration, p. 117). 



In Pteris cretica and P. fldbellata the stomata develop as follows : A 

 young epidermal cell is divided by a curved partition into two cells, one of 

 which (Fig. 61) is called the initial cell of the stoma {i.c). This is again 



