a slightly raised narrow border surrounding the diseased parts, while 

 just outside this the cuticle retains its normal healthy color, but ap- 

 pears slightly wrinkled, owing to the collapsed condition of the tissues 

 beneath. Sections through a rotten tomato at this stage show that 

 the black discolorations extend deeply into the tissues, the depth 

 depending somewhat on the size of the spot. As the malady pro- 

 gresses, the diseased parts become hard and leathery, the surface 

 assumes a greenish black velvety appearance, and finally the entire 

 fruit becomes dried and shriveled." 



Galloway identified two species of fungi in connection with the 

 spot, viz.: Macrosporium to?nato, Cook, and Fusarium so/am', Mart., 

 both of which he describes in detail. He finds the spores of the former 

 exceedingly variable, the hyphae at times long and nodulous and bearing 

 spores which cannot be distinguished from those of the genus Clado- 

 sporium. Galloway finds that the Macrosporium grows mostly in a 

 felty mass near the surface, while the Fusarium shows a much greater 

 tendency to penetrate the sound tissues. 



The results of Galloway's infection experiments are summed up as 

 follows: (i) Neither the Fusarium nor the Macrosporium has the 

 power of penetrating the sound cuticle of the tomato. (2) The 

 Macrosporium spores, when brought in contact with the exposed 

 tissue of either green or ripe fruit, produce the rot in a very short 

 time. (3) The Fusarium will grow only in fully ripe tissues or 

 tissues which have been partly disorganized through other agents. 



From the publication of Galloway to that of Jones and Grout 3 in 

 1897, tomato rot has been often mentioned in experiment station 

 bulletins, and Macrosporium tomato, or by error M. so/am, has been 

 given as the cause. A study of the so-called Macrosporium sotani of 

 potatoes by Jones and Grout resulted in the identification of two 

 species, one an active parasite and one saprophytic. Both species 

 were proved to be Alternaria. " When the study of the tomato rot 

 was taken up (by the same workers) it was found that the fungus 

 which causes the black patches on the rotting fruit was an Alternaria, 

 not distinguishable from the one on onion and potato. It was 

 further proved that this fungus did not cause the rot, for green 

 tomatoes inoculated with spores from a pure culture from the tomato 

 remained fourteen days in a moist chamber until they ripened, with- 



