D. 



individuals of both species are present, and 

 plants which are intermediate in this regard can 

 be found. Only a few populations were found to 

 have plants with stems which are conspicuously 

 villous in addition to being glandular; these 

 sites include Barber Creek (051) and Holland Lake 

 (001) in Missoula County. In Montana, the species 

 most often possesses stem vestiture which is only 

 glandular in nature. 



Foliar characteristics (i.e., leaf margin and 

 outline, degree of clasping of the leaf bases) 

 are emphasized in technical keys (Dorn, 198-4; 

 Cronquist, 1955). However, these features were 

 not found to be useful or necessary in 

 distinguishing 6. howel 1 i i and G. squarrosa in 

 the area covered during this study. 



Identifying characteristics of material which is 



in interstate or international commerce or trade: 



No interstate or international commerce or trade 

 k no wn . 



E. Photographs and line drawings: Figure 1 provides 

 a copy of the illustration of this species, 

 enlarged from Cronquist (1955). The color slides 

 (p. 7) are duplicates of those taken at the sites 

 indicated. Additional slides from other locations 

 are housed at the MTNHP office in Helena, MT . 



*» - Significance. 



A. Natural: Gr i ndel ia howel 1 i i is a fairly well- 

 marked species which may be considered a regional 

 endemic, with a bipartite distribution in northern 

 Idaho and western Montana. It is apparently 

 closely related to G. squarrosa , and thus is of 

 evolutionary significance in studies addressing 

 divergence in the genus. Gr indel ia howel 1 i i does 

 not appear to have any peculiar adaptations or 

 structures, or roles in stabilizing landforms. 

 Obligate relationships with other species are 

 unknown . 



B. Human: Gr indel i a howel 1 i i would be of scientific 

 significance in studies of the b iosystemat l c 

 relationships of the related taxa in the genus. 

 Otherwise, the species has no known agricultural, 

 economic, horticultural, or other known human 

 uses or significance. 



5. Geographical distribution. 



A. Geographical range: Gr indel ia howel 1 i i is known, 

 historically and currently, from two geographical 

 areas: a.) Benewah County, Idaho, U.S.A., and 



