400 THE WHEAT PLANT 



The characteristic four-sided ear of turgidum was absent. 



The glumes were chiefly red with some of bluish tint suggestive of 

 the turgidum grandparent ; a few forms with white chaff appeared. 



He also crossed Rivet with a " Squarehead " white-glumed wheat, 

 obtaining a beardless l with pubescent chaff of several shades of bluish 

 black. 



The F 2 exhibited a large variety of forms, some of which were quite 

 sterile. 



ii. T. vulgare, var. albidum 9 (Chidham d'automne) x T. turgidum, 

 var. dinurum (Roux velu de Beauce). 



iii. T. vulgare, var. pyrothrix ? (B16 seigle) x T. turgidum <3 (Poulard 

 Ble Buisson). Vilmorin made these crosses in 1879. The F 1 in both 

 cases was " beardless " like the vulgare parents, with short awns at the 

 tip of the ear, the glumes slightly downy and somewhat inflated like those 

 of T. turgidum. 



In the F 2 generation appeared an extraordinary series of vulgare and 

 turgidum forms tall, dwarf, bearded and beardless, some with lax ears, 

 others with very dense ears. In addition, from the hybrid (ii.) was obtained 

 a plant with the pith, glumes, and long, pointed, flinty grain of T. durum, 

 while among the progeny of hybrid (iii.) were forms of T. Spelta, some of 

 them with branched ears. 



6. HYBRIDS OF T. vulgare WITH OTHER WHEATS. 



a. T. vulgare x T. monococcum (see p. 392). 



b. T. vulgare x T. durum (see p. 394). 



c. T. vulgare x T. polonicum (see p. 398). 



d. T. vulgare x T. turgidum (see p. 399). 



e. T. vulgare x T. compactum (see p. 403). 

 /. T. vulgare x T. Spelta. 



The above crosses give fertile reciprocal hybrids, but according to 

 Stoll success is most easily secured when T. vulgare is the mother. 



i. T. Spelta, var. album $ (White Beardless Spelt) x T. vulgare, var. 

 ferrugineum $ (Bearded Red Chaff). In 1876 Rimpau made this cross, 

 the F x bearing almost beardless Spelt-like ears with slightly reddish chaff. 



The F 2 in 1878 gave all the eight possible combination forms of the 

 three pairs of allelomorphic characters (bearded and beardless, red chaff 

 and white chaff, tightly fitting and loose chaff) except white beardless 

 T. vulgare. 



In the F 3 all eight forms were found, together with a white, dense- 

 eared compactum form. 



ii. T. Spelta, var. Duhamelianum (Rose imberbe), Beardless, Red 

 Chaff x T. vulgare, var. anglicum (Blanc k duvet), Bearded, White Velvet 

 Chaff. Vilmorin records the crossing of these in 1878. 



The Fj was intermediate, the ears reddish, pubescent, with somewhat 



