YOU 2] Milliken. Calif ornian Polemoniaceae. 3 



L. Matthewsii, L. Schottii and L. setosissima treated in "Pitto- 

 nia," III (1898), and Microsteris, within which he groups several 

 species which are considered in this paper and are ordinarily 

 accepted as Gilia gracilis, treated in "Pittonia," III. "The 

 Phloxes of Western North America" (1899), is the -title of a 

 paper by Elias Nelson, in which the genus Phlox is revised. 



There is need at the present time of a general survey of the 

 order to bring together the results of the latest research, and 

 in attempting this, I have made such omissions, additions and 

 changes as seem necessary from my present point of view. 

 The material studied is that of the Herbarium of the University 

 of California, the Herbarium of the California Academy of Sci- 

 ences, and also various collections made by Dr. W. L. Jepson, 

 H. M. Hall, H. P. Chandler, J. P. Tracy, M. S. Baker, G. B. 

 Grant, and fresh material obtained during the summer of 1902 

 and 1903. 



The genera of the order, excluding those which have no 

 representative species in California, are six: Polemonium, 

 L., Collomia, Nutt., Navarretia, R. & P., Gilia, R. & P., 

 Linanthus, Benth., and Phlox, L. Several of the characters 

 formerly relied upon in denning these genera have been found to 

 separate species evidently very nearly related and vice versa. 

 Thus, Gray was led to drop the thoroughly good genus Collomia 

 and merge it in Gilia. His judgment would be justified, since 

 the characters chiefly emphasized proved unreliable, if it had not 

 been possible to find other constant characters to replace them, 

 as is shown below. The mucilagenous seeds, characteristic of 

 Collomia, are possessed by many species in all other respects 

 typical of another genus. Examples of this are Gilia aggre- 

 gata and Gilia filiformis, Linanthus Bigelovii, and many Navar- 

 retia species. The declined stamens, and appendaged filaments of 

 Polemonium are also found outside of that genus. For instance, 

 Linanthus liniflorus possesses hairy appendaged filaments, and 

 the stamens of Gilia Matthewsii are conspicuously declined. 



As is stated by Greene, in "Pittonia," I, the form of the corolla, 

 the insertion and direction of the stamens, the character and 

 number of the seeds, are all found to be unreliable as primary 

 generic characters. Greene selects as of first value the calyx. 



