SCHIZOSACCHAROMYCES FORMOSENSIS 



203 



short (about 1 to 2 /z in length) containg a nucleus and metachromatic 

 granules. The cells are often grouped in twos. (Fig. 80, 1 to 4.) 



SCHIZOSACCHAROMYCES FORMOSENSIS. Nakazawa * 



This species was isolated recently by Nakazawa from sugar prod- 

 ucts in Formosa. On beer wort, the cells are ellipsoidal or irregular 



Fig. 80. Sch. Aphalarae Calthae. 



1 and 2, Vegetative cells; 3, two 

 gametes about to unite (?); 4, egg 

 resulting from the fusion of the two 

 gametes (?); 5, 6, cells dividing by 

 budding; 7-9, cells dividing by parti- 

 tion; 10-12, beginning of germination 

 of the ascospores (after Sulc). 



Fig. 80-A. Schizosaccharomyces For" 

 mosensis, var. Tapaniensis. 



b, Copulation; d, ascs; 



; /, 

 r Na 



f 1 , germination of 



in shape (9.2-16.8 X 4.8 /z but usually about 7.2 X 6-9 /*). The 

 optimum temperature for budding in beer wort is 32 C. The ascs 

 are formed at the end of 7 days and are derived from an isogamic 

 copulation similar to those in Schizosac- 

 charomyces Pombe. They contain ellipsoi- 

 dal ascospores with no glycogen, but 

 their walls are impregnated with starch. 

 On wort, a ring is formed at 25 to 37 C. 

 and at 32 C., but not below. At 25 to 

 27 C., a scum is formed. The temperature 

 limits for scum formation are 25 to 27 C. 

 and 37 C. This yeast ferments dextrose, 



inuline, dextrine, d-mannose, d-galactose, Fig 8 'o_ B- _ Schizosaccharomyces 

 d-fructose, saccharose, maltose, raffinose, Formosensis. Vegetative Cells, 

 and a-methylglucoside. Fo of mat spore s f %?' "SiSS^^ 



Nakazawa described two other varieties J e f t | ^S &^"*^**'' 

 of this yeast, Schizosaccharomyces 



Formosensis and Schizosaccharomyces Formensis, var. akoensis. This 

 latter variety differs from the former by larger cells and larger 

 ascospores and also that the ascs are formed at the end of 5 days. 

 No ring is produced at 32 C. and scums are not produced at all. 

 Schizosaccharomyces Formosensis, var. tapaniensis, another variety, 



1 Nakazawa, R. Ueber Gamngsmikroorganismen von Formosa II. Ber. der 

 Inst. fur Exper. Forshungen, 1914. 



