ENDOMYCES CAPSULARIS 367 



vegetate, however, between 15 and 40 C. It induces a fermentation 

 in beer wort very slowly and gives off an ethereal odor. It acts en- 

 ergetically on saccharose, maltose, dextrose, levulose, mannose, galac- 

 tose, raffinose, dextrine, xylose and arabinose and feebly on rhamnose 

 and a-methylglucoside. 



Endomyces Hordei resembles Endomyces fibuliger very closely. It 

 is also related to Endomyces Lindneri. The formation of the asc by 

 simple budding, without any trace of sexuality, is responsible for this 

 resemblance. 



ENDOMYCES CAPSULARIS. Guilliermond 



Syn.: SACCHAROMYCES CAPSULARIS. Schionning l 



This yeast was described in 1903 by Klocker who isolated it from 

 pastureland in the Swiss Alps. Guilliermond finally subjected it to 

 careful study. 



Endomyces capsularis for the most part has cells in the mycelial 

 and the yeast form at the same time. It vegetates at the bot- 

 tom of the medium as a sediment or in the form of a scum. The 

 mycelium is especially well developed in the 

 scums or on solid media. It is branched, 

 with cross walls, and according to the in- 

 vestigations of Guilliermond, always has a 

 single nucleus. It is able to present dif- 

 ferent appearances. ' Some of the filaments 

 remain sterile while others form by lateral 

 and terminal budding numerous yeast-like 

 cells. Rarely there are others which form 

 small septa, dividing the thread into units 

 which break off like oidia. 



The yeast-like bodies develop especially 

 in growths of sediment. They look like 

 true Saccharomyces; their form is ellipsoidal spores (after Schi nnin g)- 

 or oval like Saccharomyces Pastorianus or Saccharomyces ellipsoideus 

 (Fig. 156 d). Many among them have a point at one or both ends. 

 They never have but a single nucleus. Aside from the yeast-like struc- 

 tures, one may find some elongated or walled cells which represent 

 yeasts in the process of making up a mycelium. 



The optimum temperature for vegetation is situated between 

 25 and 28 C. The maximum temperature is 38.5 C. and the mini- 

 mum about 0.5 C. The ascs appear under the same conditions which 



1 Schionning, H. Nouveau genre de la famille des Sacch. Comp. Rend, du 

 lab. de Carlsberg. Vol. 6, Book II, 1904. 



