690 



PHYSIOLOGY 



the diagram (Fig. 336). In a dog treated in this way it is found that the 

 amount of juice secreted by the small stomach always bears the same ratio 

 to the amount secreted by the large stomach, while the digestive power of the 

 juice obtained from the small stomach is equal to that obtained from the 

 large. This is shown in the following Table : * 



SECRETION FROM GASTRIC FISTULA AFTER SHAM MEAL 



FIG. 336. Diagram to show Pawlow's method of making a cul-de-sac of the 



cardiac end of the stomach, with vascular and nerve supply intact. 

 In A the line of the incision into the stomach wall is shown. B represents 

 the operation as completed. In A ; 0, oesophagus ; R.v, L.v, right and left 

 vagus nerves ; P, pylorus ; C, cardiac portion of stomach ; A, B, line of 

 incision. In B : V, main portion of stomach ; S, cardiac cul-de-sac ; A, 

 abdominal wall ; e, e, mucous membrane reflected to form diaphragm between 

 the two cavities. 



In this case a fistulous opening had been established into the large 

 stomach, so that the juice could be obtained simultaneously from both 

 sections of this organ. Secretion was excited by a sham meal, in which the 

 food taken by the animal dropped out of an opening in the neck, and was not 

 allowed to reach the stomach. It will be seen that the secretions in the two 

 sections of the stomach run parallel to one another, while there is an almost 

 exact equivalence between the strengths of the juices obtained from each 



* Pawlow, " The Work of the Digestive Glands " (translated by W. H. Thompson, 

 M.D.), p. 80. 



t The strength of the juice was determined by measuring the number of millimetres 

 of coagulated egg-white (in Mett's tubes) which were digested in eight hours. 



