INTKODUCTION. 9 



central one, was female and yielded seeds, as in the 

 umbelliferous genus Echinophora. There is nothing 

 surprising in the central flower tending to retain its 

 former condition longer than the others ; for when ir- 

 regular flowers become regular or peloric, they are apt 

 to be central ; and such peloric flowers apparently owe 

 their origin either to arrested development that is, to 

 the preservation of an early stage of development or 

 to reversion. Central and perfectly developed flowers 

 in not a few plants in their normal condition (for in- 

 stance, the common Hue and Adoxa) differ slightly in 

 structure, as in the number of the parts, from the other 

 flowers on the same plant. All such cases seem con- 

 nected with the fact of the bud which stands at the 

 end of the shoot being better nourished than the 

 others, as it receives the most sap.* 



The cases hitherto mentioned relate to hermaphro- 

 dite species which bear differently constructed flowers ; 

 but there are some plants that produce differently 

 formed seeds, of which Dr. Kuhn has given a list.f 

 With the Umbelliferee and Compositse, the flowers that 

 produce these seeds likewise differ, and the differences 

 in the structure of the seeds are of a very important 

 nature. The causes which have led to differences in 

 the seeds on the same plant are not known ; and it is 

 very doubtful whether they subserve any special end. 



We now come to our second Class, that of monoecious 

 species, or those which have their sexes separated but 

 borne on the same plant. The flowers necessarily 

 differ, but when those of one sex include rudiments 



* This whole subject, including Domestication,' chap. xxvi. 2nd 



pelorism, has been discussed, and edit. vol. ii. p. 338. 

 references given, in my * Variation t ' Bot. Zcitung,' 1867, p. 67 

 of Animals and Plants under 



