170 THE MASTIGOPHORA 



forming, as products of metabolism, glycogen-like bodies (Klebs). H. 

 muris is found in the intestine of mice (Wenyon). Lamblia intestinalis 

 is found in the intestines of various mammals, and is not infrequently 

 parasitic in man. It is not thought to be pathogenic. Spironema, Klebs. 

 Polymastigote ; flagella arising in pairs at the margins of the spiral 

 mouth-grooves. 



SUB-TRIBE 4. LOPHOMONADINA, exclusively parasitic in the rectum of 

 insects. This sub-tribe is regarded by some authors as having closer 

 affinities with the Ciliata. It has already been described under the 

 heading Family Trichonymphidae in Fasc. II. p. 417 of this Treatise. 

 The genus Maupasia (Schewiakoff) has the anterior part of the body 

 covered with cilia, but at the posterior end it bears a long flagellum. By 

 some authors it is regarded as a Polymastigine flagellate, but its affinities 

 seem to be with the Ciliata. Freshwater. Hawaii. Monomastix, Roux, is 

 another genus with a polar flagellum and cilia in longitudinal rows. 

 There are said to be two meganuclei and two micronuclei. This genus 

 should also be included in the Ciliata. 



ORDER 2. Euglenoidea. 



The second order of Lissoflagellata comprises the most highly 

 organised members of the sub-class. This high degree of special- 

 isation is indicated by the structure of the pharyngeal armature of 

 the tribe Peranemina, which consists of two converging rods, which 

 can be protruded from the base of the oral funnel. 



With regard to the nutrition of Mastigophora as a class, to 

 which allusion has already been made, it is necessary, even from a 

 purely systematic standpoint, to consider (1) the nature of the 

 food ; (2) the mechanism of ingestion ; (3) the products of meta- 

 bolism. In holozoic nutrition the food consists of bacteria, other 

 monads, swarm-spores of Algae, starch, and the like. The modes 

 of ingestion by which these food-bodies are conveyed into the pro- 

 toplast of the feeding organism are of five principal kinds, namely, 

 pseudopodial ingestion (Pantostomatina) ; vacuolar ingestion (Mono- 

 mastigoda, Paramastigoda, Isomastigoda, Choanoflagellata) ; rostral 

 or suctorial ingestion (Heteromastigoda) ; stomatic ingestion, by 

 which the food sinks into the protoplasm through one (Monostoma- 

 tina) or two (Distomatina) points of least resistance situated in one 

 or two depressions (oral grooves) below the insertion of the flagella ; 

 pharyngeal ingestion (Peranemina). 



Not only does the mode of feeding distinguish the Peranemina 

 from all other Flagellata, but they are further distinguished by 

 their well-marked, spirally striated periplast or cuticula. In the 

 Euglenoidea the periplast is generally a striated, resistent proteid- 

 membrane. 



The vacuole-system of the Euglenoidea consists of a non-con- 

 tractile or feebly contractile reservoir provided with an excurrent 



