REPORT ON THE SLUGS. 433 



outer side of the kidney is the lung, which also proceeds very far backwards, extending 

 to a point just beyond the terminal portion of the kidney (PI. XL. Fig. 9). The 

 terminal portion of the respiratory duct opens separately from that of the combined 

 rectum and ureter, and slightly above it, so that there is a distinct cloacal chamber 

 into which these two ducts open (PI. XL. Fig. 11). The external opening of this 

 chamber lies on the right side of the hypnotum (PL XL. Fig. 2). 



IV. TJie Generative System. 



Although conforming in general to the type of reproductive organs common to 

 Veronicella, there are certain well-marked features which have specific value. 



The hermaphrodite gland is pyriform in shape, and has a closely convoluted duct 

 which is enclosed in a fine membranous sheath (PI. XL. Fig. 12, h. d.) at its base, 

 and lying upon, and in life completely hidden by, the albumen gland, is a small 

 diverticulum, the vesicula seminalis (PI. XL. Fig. 12, v. s.). The albumen gland is 

 a loose glandular organ not unlike that in V. leydigi, Simr. The oviduct is a long 

 wide tube, twisted many times upon itself. The middle portion is the widest, while 

 its commencement is the narrowest ; it also becomes narrow again before opening 

 externally. The vas deferens is a narrow tube, and bears at its lower end a large 

 spermatocyst, which has a duct of some considerable length (PI. XL. Fig. 12, sp.). 

 At the opposite side of the vas deferens, at the point where the duct of the 

 spermatocyst enters, a duct is given off to the receptaculum seminis. The form and 

 position of the spermatocyst, together with its duct, is very characteristic of this 

 species and separates it from any allies. The receptac^dum seminis is a large ovoid 

 body, with a long tivisted duct (PL XL. Fig. 12, rs., rd.). 



In the male organs I follow Simroth in terming the penis-gland of Semper, the 

 dart-gland or dart-sac. 



The penis is enclosed in a thick muscular sac, and in general outline and structure 

 (PL XL. Fig. 13, p.) agrees very closely with that figured by Simroth for V. lei/digi\ 

 being long and conical in shape and having an oblique slit at the top; there is, 

 however, in V. willeyi, quite a large vestibule into which the penis opens, slightly 

 above the opening of the dart-sac. Attached to the penis, just below the point of 

 origin of the vas deferens, is a strong retractor muscle, and a smaller one just below 

 the vestibule (PI. XL. Fig. 13, rm., rm.'). The dart-sac is rather longer and \\ider 

 than the penis (PL XL. Fig. 13, ds.). At the base of the sac there is a small 

 conical papilla, the dart (PL XL. Fig. 13, d.), and connected with this, but outside 

 the actual dart-sac, are a series of long thread-like accessory glands (PL XL. Fig. 

 13, ac. gl.) ; these all open at the base of the papilla or dart, the opening at the 

 apex of the dart forming a common orifice. The number of these accessory glands was 

 not constant, the average is 15, but in one case there were only 12, while in another 

 there were as many as 20. 



Whether or not this dart-gland is homologous with the dart-sac in other 

 Pulmonates, e.g. Parmarion, Bamayantia, or with the dart-sacs in certain species of 

 Helix and Zonites, which contains an imperforate calcareous dart, I at present hesitate 



' Cp. T. c. Tiif. L, Fig. 4. 



