536 A DESCRIPTION OF THE ENTOZOA COLLECTED BY DR WILLEY 



thinner though they retain their muscular character. The lining of this part of the 

 penis usually shows a frayed margin something like a ciliated surface but I doubt" 

 if the appearance is due to cilia. The lumen in many sections is semi-lunar shaped, 

 the presence of a typhlosole reducing it to a curved slit (Fig. 5, Plate LIV.). All 

 along the outermost portion of the penis the tube receives the secretion of numerous 

 glandular cells more or less aggregated into a gland. The penis curves to the left 

 of the "egg-cavity" and opens into a very shallow small genital sinus a little to the 

 left of the middle line not very far behind the posterior limit of the pharynx (Fig. A 

 in text). 



Female. The female organs are equally complicated. The ovary is single, dorsal, 

 posterior and slightly to the left of the middle line. It consists of compact cells, the 

 ova ; the less mature show a very well marked chromatin tangle in the middle of their 

 nuclei. The ripe ova have that curious cubical look with rounded corners which is 

 characteristic of many Trematode eggs. The oviduct is short and opens almost directly 

 into the shell-gland complex. 



The yolk glands are densely staining compact glands lying ventral to the testes, 

 stretching in front of that organ for about a third of the way or perhaps one half 

 the way along the body. They consist of cells apparently crowded with yolk granules. 

 Their ducts, one on each side, converge and enter the shell-gland complex (Fig. A in 

 text). 



The shell-gland is median, it consists of numerous glandular cells surrounding and 

 for the great part forming the walls of the structure which I have referred to as 

 the shell-gland complex. I have not succeeded in tracing out all the details of this 

 minute organ. I have traced the oviducts, and the ducts of the yolk glands into a 

 cellular complex with a canal. The canal is I believe continuous with the very short 

 oviduct and its cellular walls the shell-glands. From this I have traced the beginning 

 of the uterus with young eggs in it, but I can offer no explanation as to how the small 

 ovum with its large nucleus is packed up with the yolk granules and each surrounded 

 by its cuticular egg-shell. It must however all take place in this minute organ. 



The lower end of the oviduct soon widens until we find a tube with thin walls 

 and a uniform lumen which passes from side to side of the body forming some thirty 

 loops (Fig. 5, Plate LIV.). These are very much simplified in the diagi-am (Fig. A 

 in text). 



Towards the anterior end of the body this oviduct opens into an irregular space 

 which I have called the " egg-cavity." Its walls stain deeply and when I first saw 

 it in section I took it for a part of the ventral hollow which projects so far into 

 the under.side of the body. It however has no communication with the exterior except 

 through the vagina. The latter leaves its anterior end and is at first a fairly wide 

 tube. During its course two similar and highly peculiar organs open into it, one nearer 

 the inner and the other nearer the outer opening. 



Each of these organs has a somewhat Hattened spherical shape, one surface pro- 

 jecting freely into the lumen of the vagina. The substance of the organ is homogeneous, 

 no cells can be detected in it and the whole stains lightly but uniformly. Scattered 



