S. Ikkno r)3 



be asked, whether the Oenoiheni -hik^ch^h U8e<i by the»t» iiuthorH hiul 

 genotypic constitutioiiH exactly similar to thns(» wnvil by the Dutch 

 botaninL If thi» be n^ally true, then this hybri<liwiti<ni stuMiiM t^» have 

 mm\e remMiibhince to what I have seen in .S'a/i.r-crnHH unHer ennMi«h»- 

 ration. 



My rc»8ult«(»n the hybridisjition Salir inuld'ncrvis x Sulix gnu^lUUfUi 

 ap-ee with those reconled by Lidt'orss fur liiibns, inasmuch us 1 havi* 

 also obUiiniMl lv>th genuine hybrids jis well us plants uf purely mat<*rnal 

 ty|K'. The (iiflferenee between the tw(> hybridisiitions li«*s oidy in this: 

 while Lidfoi^vs obtained l>oth real and false hybrids in one and the same 

 year. I was aWe to obtain them in diti'erent years (1910 and 1911). 

 That the dilTerence of results in thwso two yeai-s is due to the ^'rnolypic 

 differences of male and female plants used by me may be absolutely 

 denietl, for. jvs above stated, the female plant used in both civses w.us 

 one and the Siime tree, and the male plant in 191 1 was either the same 

 with, or at least derived in a vegetative way from, that used in 1910. 

 As I obtained ditlerent results in ditferent years, one might be dis- 

 j)osed to think that whether the ot!*spring will be real hybrids or of 

 purely maternal tyi)e is dejK'udent on external conditions, which indeed 

 may be true. But I think it ecpially likely that our plant contains 

 two kinds of eggs, giving genuine and false hybrids, respectively, 

 just as we have in Thalictrum purpurascens^ and some species of 

 Hieracium^ two distinct kinds of eggs, i.e. those .which can develop 

 only after having been fertilised, and those which are able to develop 

 parthenogeneticiilly, though the final decision of this (piestion will be 

 of course impossible without performing further breeding experiments. 



Now some remarks about the sex of false hybrids. Rubus is always 

 hermaphrodite, so that both real and false hybrids are naturally of this 

 nature. The sex of those of Melandrium obtained by Gartner is 

 unknown, for he has stated nothing about it^ Those of Fragaria 

 virginiana % x F. elatior J^ were either male <a- female*. Those of Salix 

 got by me, almost fifty in number, were female without a single excep- 

 tion, so that they may be well siiid to be of maternal type in the most 

 strict serine of the luord, for even the sex has been inherited. Further- 

 more, that our false hybrids will breed true in later generations like 



1 Overton, Hot. (iaz. Vol. xxxiii. l'.K)2, pp. 3G3— 374 ; lU-r. <L heutxch. Hot. Ge*. 

 Bd xxu. 1904, pp. 274—283. 



* Ostenfeld, IloUiuhk Ti(hfkrift, Bd xxvii. l'.*(»6, pp. 221 -J.'iO ; Xeitx. f. itnl. Ahst,im- 

 mungit' u. Vererhnunslehre, Bd in. 1910, pp. 241—28^). 



» Lx. * SoliiiH-Laubacli, / c. p. o3. 



