S. r IIaki.am) 105 



He ri'ffrs U* Forms a — c iih onliiiary vyo or .smiill ryr. Koriiis e 

 1111(1 /* art' tn'rtU'd iw onr mulcr the imiiu' ll«»l.st«'in, ihun tin* (roloiir 

 (iiilUTn of 11 viirioty hiivini; that namr. Korni r/ is cjillrd bir^a- eye, and 

 is eoiiMidoriHi i\s xhv lictfrozyi^nt*' Ix'twrin H<»lst-4Mii ami Small ^y(^ 

 Form ff n'pnrsontss a k^ciutically ditTt'iciit ty|H' of rye In it thr |»i^- 

 mcnUHJ aiiNi surnmrnls l\w hilum, Imt thr micrnpylar rnd nf ihr Hrua 

 has the marj^in vt-ry indistinct ; fine <l<»ts of |Mt^mrnt, extend i»v(i- tin- 

 micropylar cnti of th*- scrd. Spillnian cmIIs this latt«i- t.y|M' tin- Watson 

 oyi", from a variety rhanic'tcrisrd hy this ty|M' of |».ill«rn. 



Ho investigat.i*d the results of 21 crosses iMtweeii Small eye and 

 Solirl colour. In all crust's the Ft was Solid coloui-. In the F.. he 

 oht^iined the following types: Solid, Watson, Holstein, Lirge eye, und 

 Small eye. The ratio between them approximaU'd to !>:*}: I : 2 : I. 



He wivs able t^ fonnulate four hypotheses to exj)lain his ratios. A 

 summary may hv given of the hypothesis which he consi<lers as best 

 adaptrcd to explain his results. 



The behaviour of the cross Small eye by Solid colour indicates that 

 these two ty|>es ditTer in two factors which are transmitted inde|x;ndently 

 of each other. Acconlingly the Watson pattern may be regarded as due 

 to a fiict^r W, and the Holstein pittern iis due to a factor H. The 

 formulae* of the tyj)es appearing in F. will be: 



1. Solid WWHH 



2. Watson WWhh 

 8. Holstein wwHH 

 4. Small eye wwhh. 



The present writer has investigated in detail the results of three 

 crosses between Small eye and Solid colour. In inj cjise have such simple 

 results as those of Spillman been obtained, but the data secured have 

 provided an indirect confirmation of the latter's work. An account of 

 the results obtained in the present work will now be given. 



The Experimental Results. 



Cross 1. Black eye) , jRouncevaP. 

 Small eyej (Solid colour. 



Several crosses were made, and the F^ was in all cases Solid colour. 



The F, 



The results are j>resented in Table VII. The following lyi)es ap- 

 peiired : Soli<l, Watson, Holstein types e and /, Large eye spotted, and 



' For description .see p. 114. 



