372 



THE ARACIINOIDAE. 



§ 301. 



ganiiuo 



The ventral chain of the Pjcnogonidae is composed likewise of four 

 ganglia, but these, which send off, each, a nerve from its side to the corre- 

 sponding foot, are contiguous, and the first connects with the ovoid cerebral 

 11 by two lateral commissures.'^* 



With the Ar;ineae, the central portion of the nervous system consists of 

 a large sub-ocsophageal ganglion, and another, smaller and above the oesoph- 

 agus. They are separated from each other only by a narrow fissure 

 through which the oesophagus passes. The super-oesophageal ganglion, 

 which is somewhat emarginated in front, corresponds to the brain, and 

 sends off nerves to the eyes and cheliceres. The sub-oesophageal gan- 

 glion, situated in the middle of the cephalothorax, sends off, on each side, 

 four larger processe;^, from which arise the nerves of the feet. Its anterior 

 border supplies, moreover, the nerves of the two palpi ; and from its poste- 

 rior margin pass off two nerves for the abdominal viscera.'"'' 



The nervous system of Galeodes,"'^ Pkrynus, and Thelyphomis,^^ has a 

 like disposition. 



The central mass of the nervous system of the Phalangidac begins by 

 twC conical, contiguous cerebral ganglia, which connect with a sub- 

 oesophageal, fused ganglion, by two short lateral commissures. This ven- 

 tral ganglion is composed of a transverse portion, which is situated in the 

 centre of the cephalothorax, and of two lateral portions which consist, each, 

 of an anterior or larger, and a posterior or smaller lobe. These lobes send 

 oft" nerves to the eight legs, and in front, others to the palpi ; while from 

 the posterior border of the transverse portion pass off several nerves to the 

 viscera of the abdomen.® * 



With the Scorpionidae, the nervous system is very highly developed. 

 The brain"", which is not large, is composed of two spheroidal, super- 

 oesophageal ganglia fused together. Above, and in front, they send off 

 nerves to the eyes and the cheliceres ; and below, they connect with the 

 first ventral ganglion by two short, large filaments, which embrace the 

 oesophagus. The first ventral ganglion is pretty large, being the result, 

 probably, of the fusion of several ganglia. It is situated in the middle of 

 the cephalothorax, and sends nerves to the palpi and to the eight legs. In 

 the rest of the body there are three ventral ganglia, smaller, and followed 

 by four others situated in the tail. All these ganglia are connected by 

 double, longitudinal commissures, and the posterior seven give off, from 

 each side, two nerves ; while from the last ganglion arise also two others, 

 which, passing backwards, soon unite and extend to the very extremity of 

 the tail, sending off" nerves right and left,® 



4 Quatrf/agex, loc. cit., 77, PI. I. fig. l:* 2^; also 

 PI. 11. fig. 2, 3 {Ammothea and Phoxicliilus). 



5 Treviranux, Ueber d. inn. Bau d. Aiach. p. 

 U,Taf.V. fit,'.45,andZeitsoli. f. Physiol. IV. p. 94, 

 Taf. VI. fig. i ; Lyonet, loc. cit. p. 4U5, PI. XXI. 

 tig. 22 ; Brandt, Mediz. Zool. II. p. 90, Taf. XV. 

 fig. 3, 4, or Ann. d. Sc. Nat. Xlll. p. 184, PI. IV. 

 tig. 4. Dugis, Ibid. VI. p. 174; Grube, loc. cit. 

 ]). 302, and finally Owen, Lectures, &c., p. 2.5.5, fig. 

 109. Tills last autlior has roprusenfjd, in a very 

 Instructive uiainicr, the nervous system of a My gale 

 seen in protile. 



fi HIannhnrd, loc. cit. p. 1384. 

 <■ Van de.r Hoeven, Tijdschrift. loc. cit. IX. 

 1842, p. OS, and X. 1843, p. 30'J. 



8 The nervous system had already been partially 

 described by Treviranus (Verm. Schrift. I. p. 38, 

 Taf. IV. tig. 24) ; but especially, and with full de- 

 tails, by Talk, loc. cit. p. 324, PI. V. fig. 31. 



9 For the nervous system of the Scorpionidae, 

 see Treviranus (Bau. d. Arach. p. 14, Taf. I. tig. 

 13, and Zeitschrift f. Physiol. IV. p. 89, Taf. VI. 

 fig. 1-3, and Mailer, loc. cit. p. 60, Taf. I. fig. 5, 

 7) ; but especially Newport''!) excellent description 

 (I'hilos. Trans. lS4;j, p. 200, PI. XII.) ; he has 

 traced, with Androctonu.i, the nerves of the e.\- 

 tremities even into the tjirsal articles and terminal 

 hooks.* 



*[§ 301, note 9.] See also Dufour (Ann d. behind the thoracic mass, from which passes 

 .Sc. Nat. XV. 1851, p. 250). This anatomist has off a pair of nerves to the pulmonary organs 

 found a fourth abdominal ganglion, situated just (JScorpio occitanus). — Ed. 



