PALAEONTOLOGY 



species now known as Morosaurus brevis. Still smaller vertebra of the 

 same, together with teeth erroneously assigned by Mantell to the under- 

 mentioned Hylceosaurus, from Cuckfield and Tilgate Forest, have been 

 made the types of another species of dinosaur by the present writer 

 under the name of Pleuroccelus valdensis, the generic name being first 

 given to a North American dinosaur. 



Quite another type of dinosaur is indicated by Hylceosaurus armatus, 

 a genus and species known only from the Wealden of Battle, Bolney 

 and Tilgate. It was armed with a number of large bony spines, 

 probably carried in one or more rows along the back. A lower jaw 

 from the Wealden of Cuckfield described by Mantell under the name of 

 Regnosaurus northamptoni probably indicates a reptile nearly allied to, 

 if not identical with, the last ; the specific name was applied in honour 

 of a former Marquis of Northampton, and does not refer to the locality 

 of the specimen. The carnivorous dinosaurs are represented by a 

 species of the Jurassic genus Megalosaurus, which has been named by 

 the present writer M. oweni ; its remains have been recorded from 

 Battle, Cuckfield and Hastings. Some remains of the same genus from 

 the Wealden of the county have been assigned to a continental species, 

 M. dunkeri. Of the crocodiles of the Wealden of the county, one of 

 the most abundant is Goniopholis crassidens, a genus and species first 

 described by Mantell from the Purbeck of Swanage ; its remains occur 

 at Cuckfield and Horsham, and a nearly perfect skull is preserved in 

 the Brighton Museum. The bony plates protecting the body of this 

 crocodile are articulated together by means of a peg-and-socket arrange- 

 ment. Vertebra from Cuckfield have been regarded as indicating a 

 second species of the genus, named G. carinata. Crocodilian teeth from 

 Cuckfield and Tilgate Forest of a different shape to those of Goniopholis 

 — having a pair of sharp vertical ridges on opposite sides of the crown — 

 have been made the type of a distinct genus under the name of Sucho- 

 saurus cultridens. Finally, a more modern type of crocodile from the 

 Wealden of Cuckfield and Hastings has been described as Heterosuchus 

 valdensis ; but it does not seem certain that it is really distinct from a 

 crocodile from the Belgian Wealden named Hylaochampsa. 



Several kinds of freshwater tortoises are known from the Wealden 

 strata of the county, most of these belonging to a group now restricted 

 to the southern hemisphere. An exception in this respect is however 

 a very remarkable Chelonian from Cuckfield and Tilgate Forest described 

 by Mantell as Trionyx bakewelli, but now known as 'Tretosternum bakewelU. 

 Although the shell is sculptured, the nearest living ally of the genus 

 (unfortunately very imperfectly known) is the American snapper 

 {Chelhydrd). A second, unnamed, species of the genus occurs in the 

 Wadhurst Clay of Hastings. The Chelone belli of Mantell, from Tilgate 

 Forest, is now referred to the genus Hylceochelys, typically from the 

 Purbeck of Swanage, where it is represented by H. latiscutata. To the 

 latter species is assigned an imperfect shell in the British Museum from 

 the Wealden of Burwash in the county under consideration. The 



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