200 PLANT BIOLOGY 



dry weather the margins of the leaves roll inward, and the leaves 

 fold closely against the stem, thus protecting the delicate assimi- 

 lating tissue. 



The antheridia and archegonia of polytrichum are borne in 

 groups at the ends of the branches on different plants (many 

 mosses bear both organs on the same branch). They are sur- 

 rounded by involucres of characteristic leaves termed perichcetia 

 or periductal leai'es. Multicellular hairs known as paraphyses are 

 scattered among the archegonia and antheridia. The involucres 

 with the organs borne within them are called receptacles, or. le^s 

 appropriately, " moss flowers." As in marchantia, the organs are 

 very minute and must be highly magnified to be studied. 



The antheridia are borne in broad cup-like receptacles on the 

 antheridial plants (Fig. 297). They are much like the antheridia 



of marchantia, but they stand free 

 among the paraphyses and are not 

 sunk in cavities. At maturity they 

 burst and allow the sperm cells or 

 spermatozoids to escape. In poly- 

 trichum, when the receptacles have 



fulfilled their function, the stem con- 

 Fig. 297. - Section through a tinues to grow from the center f 



receptacle of polytri- the cup (w, Fig. 295 ) . The arche- 



chum commune, showing • „ 1 „• ,( ., 



. ,. s gonia are borne in other receptacles 



paraphyses and antheridia. ,.„. , m, ,., 



on different plants. 1 hey are like 

 the archegonia of marchantia except that they stand erect on the 

 end of the branch. 



The sporogonium which develops from the fertilized egg is 

 shown in a, l>, Fig. 295. It consists of a long, brown stalk bearing 

 the spore-case at its summit. The base of the stalk is imbedded 

 in the end of the moss stem by which it is nourished. The 

 capsule is entirely inclosed by a hairy cap, the calvptra, b. The 

 calyptra is really the remnant of the archegonium, which, for a 

 time, increases in size to accommodate and protect the young 

 growing capsule. It is finally torn loose and carried up on the 

 spore-case. The mouth of the capsule is closed by a circular lid, 

 the operculum, having a conical projection at the center. 



The operculum soon drops, or it may be removed, displaying a 

 fringe of sixty-four teeth guarding the mouth of the capsule. This 

 ring of teeth is known as the peristome. In most mosses the 

 teeth exhibit peculiar hygroscopic movements ; i.e. when moist 

 they bend outwards, and upon drying curve in toward the mouth 

 of the capsule. This motion, it will be seen, serves to disperse 

 the spores gradually over a long period of time. 



Not the entire capsule is filled with spores. There are no 

 elaters, but the center of the capsule is occupied by a columnar 



