BOLTONIELLA TENUITEGMINATA. 57 



from Commentary, France, that it is difficult to refrain from classing the wing as 

 blattoid. The neuration is extensively branched and the wing-boundaries are not 

 well marked, but the wing appears to have been somewhat quadrangular in outline, 

 with a sinuous inner margin, and the base much broader than is seen in the ordinary 

 form of blattoid hind-wing. The outer margin seems to have been straight, a 

 portion of it still remaining in the middle third. The two margins merge in a 

 well-rounded apex. The subcostal area is narrow, strap-shaped, and probably 

 extended over the whole length of the outer margin. No traces of cross-nervures 

 can be seen on it. The radius divides near the base, giving rise to a series of 

 branches which curve inwards as they approach the apex of the wing. The course 

 of the branches is irregular, the interspaces widening and narrowing, possibly 

 owing to the wing having crumpled during deposition. The median divides into 

 two branches low down, each of which is repeatedly forked, the final divisions 

 becoming attenuated and untraceable before the inner margin of the wing is 

 reached. The course of the cubitus is obscured by a reed-like plant, only two 

 basal portions being distinguishable. The anal area is filled by a broad series of 

 thread-like veins, which sweep obliquely inwards in a fan-shape, and occupy a large 

 part of the inner margin. 



The inner margin of the wing to a third of its total length is quite filmy, the 

 veins crossing the area as faint shadowy lines. The distal two-thirds is more 

 strongly impressed, while in the broad base of attachment the stems of the 

 principal veins seem to have been more than usually robust. No trace of trans- 

 verse nervures or reticulation can be seen. 



Affinities. The nearest analogues to this wing are, I believe, the forms described 

 by Brongniart as Lamproptilia grand' euryi and L. stirrupi (' Etudes sur le Terrain 

 Houiller de Commentry,' vol. iii [1893], pp. 46770, pi. xxxv [19], figs. 79). 

 It appears to be more closely related to L. stirrupi than to L. grand'euryi, but is 

 more quadrangular, and its costal area is broader. The anal portion of the wing 

 is of greater tenuity, and occupies fully half of the inner margin. 



Family BRODIID^:, Handlirsch. 



1906. Handlirsch, Die Fossilen Insekten, p. 113. 



1919. Handlirsch, Revision der Paliiozoischen Insekten, p. 73. 



Wings in which the anal area is much reduced and specialised, the radius 

 undivided, and the median, cubitus, and anal veins arched and directed back 

 towards the inner margin. 



From a re-examination of the originals in the British Museum, Handlirsch has 

 concluded that the family does not belong to the Palaaodictyoptera, but to the 

 Order Megasecoptera. This view seems at variance with his own definition of the 

 8 



