NO. 5.] USTILAGINE/E OF CONNECTICUT. 29 



through gelatinization ; they are of large size, 16-35/i. Figs. 

 11, 12, 39. 



This is the type of the family, and is one of the larger 

 genera. The sorus is quite like that of Ustilago, but the 

 spores are usually considerably larger. Saccardo records 53 

 species ; 1 5 have been found in North America, but only 2 in 

 Connecticut. The important stinking smuts of wheat, which 

 cause so much damage in the western wheat districts, belong 

 in this genus ; but, as wheat is little grown in this state, neither 

 species has been reported here. The smooth-spored species, 

 Tilletia f ceteris (B. & C.) Trek, however, has been found, on 

 microscopic examination, in ground cattle food offered for 

 sale in the state, apparently made from smutted western wheat. 

 The cattle did not relish this food. 



Key to Species of Tilletia. 



I. Spores reticulate T. Anthoxanthi. 



II. Spores apparently verruculose T. Maclagaui. 



Tilletia Anthoxanthi Blytt. Figs. 12, 39. The sori arc 

 ovate, about 3 mm. in length, and usually occur in all of the 

 ovaries of the spike ; they are somewhat hidden by the envel- 

 oping glumes, and at first are covered by a thin plant mem- 

 brane, which eventually ruptures, disclosing the dusty, reddish 

 black spore mass. Hyaline cells, or immature spores, are 

 mixed with the spores, and are smaller than those, and have 

 thin to medium-thick walls. The mature spores are reddish 

 brown, ovoid to spherical, reticulate (3-6^ wide and 1-3/z 

 deep), and 24-30/*, occasionally even 34/i, in length. 



Host and Distr. : Anthoxanthum odoratum, Whitney- 

 ville, July 6, 1902. 



The sori are so inconspicuous in the ovaries that the in- 

 fected spikes scarcely differ in appearance from the normal. 

 See Fig. 39, in which the spike is magnified two diameters. 

 This European species has been found in this country only at 

 the above station. 



Tilletia Maclagani (Berk.) Clint. Fig. 11. The sori are 

 found in the ovaries, and rarely also in the anthers, and are in- 

 conspicuous, being concealed by the enveloping glumes ; upon 



