204 LINES OF DESCENT. Chap. IX. 



developed state, of Goodyera, Epipactis, and Spirantheg, 

 all members of the Neotteae, could by further slight 

 modifications have given birth to the tribe of the 

 Ol^hre*. 



Hardly any question in Natural History is more 

 vague and difficult to answer than what forms ought 

 to be considered as the highest in a large group ; * for 

 all are well adapted to their conditions of life. If we 

 look to successive modifications, with differentiation of 

 parts and consequent complexity of structure, as the 

 standard of comparison, the Ophreae and Vandeas will 

 stand the highest among the Orchideae. Are we to lay 

 much stress on the size and beauty of the flower, and 

 on the size of the whole plant ? if so, the Yandeai are 

 pre-eminent. They have, also, rather more complex 

 pollinia, with the pollen-masses often reduced to two. 

 The rostellum, on the other hand, has apparently been 

 more modified from its primordial stigmatic nature in 

 tlie OphreaB, than in the Yandea?. In the Ophres the 

 stamens of the inner whorl are almost entirely sup- 

 pressed, — the auricles — mere rudiments of rudiments — 

 being alone retained; and even these are sometimes 

 lost. These stamens, therefore, have suffered extreme 

 reduction; but can this be considered as a sign of 

 highness? I should doubt whether any member of 

 the Orchidean order has been more profoundly modified 

 in its whole structure than Bonatea speclosa, one of. the 

 Ophrea?. So again, within this same tribe, nothing 

 can be more perfect than the contrivances in Orchis 

 ivjramidalis for its fertilisation. Yet an ill-defined 

 feeling tells me to rank the magnificent Yandese as 

 the highest. AVhen wa look within this tribe at the 



* 



1 he fullest anil the most able his ' Entwickeluugs-Gesetze der 

 dispussion .*Ji this difficult .subject Orgaiiischen Welt/ 1858. 

 is hv I'rotcsaor H. (J Jiionu iu 



