Chav. i\. concluding kemarks. 291 



<iecasionally producing capsules under cultivatiois 

 likewise come under this head. 



Of species which regularly fertilise tliemselves with- 

 out any aid and yield full-sized capsules, hardly any 

 case is more striking than that of Ophrys cfpifera^ which, 

 was advanced by me in the first edition of this w ork : 

 To this case may now be added two other European 

 plants, Orcliis or Neotinea intacta and Epipactis viridi- 

 Jiora. Two North American species, Gymnadenia 

 tridentata and Platanthera hijperhorea appear to be in 

 the same predicament, but whether when self- fertilised 

 they yield a full complement of capsules containing 

 good seeds has not been ascertained. A curious 

 Epidendrum in South Brazil which bears two additional 

 anthers fertilises itself freely by their aid ; and Dendro- 

 hiiuii cretaceum has been known to produce jaerfect self- 

 fertilised seeds in a hothouse in England. Lastly, 

 Spjirantlies ausiralis and tw^o species of Thelymitra, 

 inhabitants of Australia, come under this same head. 

 No doubt other cases will hereafter be added to this 

 short list of about ten species which it appears can 

 fertilise themselves fulh^ and of about the same 

 number of species which fertilise themselves imper- 

 fectly when insects are excluded. 



It deserves especial attention that the flowers of all 

 the above-named self-fertile species still retain A'arious 

 structures which it is impossible to doubt are adapted 

 for insuring cross-fertilisation, though they are now 

 rarely or never brouglit into play. We may therefore 

 conclude that all these plants are descended from 

 species or varieties which were formerly fertilised by 

 insect-aid. jMoreover, several of tlie genera to whicli 

 these self-fertile species belong, include other species 

 which are incapable of self-fertilisation. Thelymitra 

 offers indeed the only instance known to me of two 



U 2 



