100 Inheritance in Barley 



This classification of the forms of the lateral floret, although adequate 

 for the recognised natural races of barley, cannot be infallibly applied 

 to hybrid generations, even when types (C), (D) and (E) are employed. 

 The late tillers of a single plant may differ from the early ones, and 

 upon any ear more than one type of lateral floret may occur, e.g. (C) 

 and (D), (D) and (E), etc. 



The exact morphology of (F) and (G) will have to be determined by 

 sections of develpping ears at successive early stages. 



Hordeum inerme — an awnless distichum obtained first by Rimpau 

 from the cross H. Steudelii x H. trifurcatum — offers many complexities 

 when used in crosses, in connection with the form of the natural floret. 

 Results so far obtained upon certain Fz's in which H. inerme has been 

 used, have not given absolutely consistent results, and consequently 

 they are not included in this paper. H. inerme appears to be the 

 key to the problem of inheritance of several characters, e.g. lateral floret 

 and awns. 



Details concerning the classification by form of lateral floret are to 

 be found in any systematic account of the cereals. The most practicable 

 classification appears to be that of Beaven (3). 



III. The Inheritance of the Lateral Floret. 



Observations have been made upon a considerable number of crosses 

 between 6-row and 2-row (distichum) barleys. In the notation of 

 §11 above these may be represented by (A) x (F). In some cases 

 reciprocal crosses were made, but as both directions of the cross gave 

 the same result — a fact established by Biffen (4) and (5) for many 

 barley crosses — the (/ and $ parents are not separately designated in 

 the tables of results. 



At the outset, an attempt was made to classify in the F^^ all the 

 various forms of lateral which were neither (A) nor (F). Table I 



TABLE I. Six-row x Two-row. 



