290 Colour and Markings in Pedigree Hereford Cattle 



ance of Wetmore Laurel's ancestors further than the first parental 



generation. The pedigree so far as the characters of the animals is 



known is set forth in the accompanying diagram. 



? (? 



Graceful x Lowland Paradigm 



(Vol.36, p 533) 

 DD 



(26986) 

 NW 



Gaylass 2nd 



(Vol. 46, p. 849) 



DNW 



Bounds Chance 

 (29544) 

 DNW 



Wetmore Daisy x Laureate 8th 



(Vol. 44, p. 606) 

 NW 



(29174) 

 DNW 



Gaiety x 



(Vol. 47, p. 899) 

 DNW 



Wetmore Laurel 

 (33259) 

 DNW 



Albynes Gaylad 

 (Vol. 50) 

 DD or DN 

 Diagram 3. A pedigree set forth to show how the dark-necked character, which is 

 normally dominant, may be carried for several generations by typically marked cattle, 

 its expression being inhibited by the factor for excessive white, until a suitable recom- 

 bination of the factors occurs in which that for too much white is eliminated, and the 

 dark-necked character shows again in full force. 



The matings in which the " dark-necked " character has been con- 

 cerned may be tabulated as follows : 



{Lady 4th x Newstead (30814) 

 X Bounds Chance (29544) 

 Playful X Lowland Paradigm (26986) 

 ,, X Bounds Chance (29544) 



Totals 

 DD xDNW Shelsley Cypress x Bounds Chance (29544) 



DNW X DNW Gaiety x Wetmore Laurel (33259) 



Summary : The evidence so far to hand leads one to believe that 

 the " dark-necked " factor is dominant to that for normal colouration, 

 from which it segregates in the ordinary Mendelian ratio, but it becomes 

 more or less rebessive in the presence of the factor- for excessive white, 

 which inhibits its full expression, so that a mating between outwardly 

 normal individuals may give rise to a grade -f 3 animal, as in the case 

 of Gaiety and Wetmore Laurel just cited. 



III. Pigment Surrounding the Eyes. 



"Red-eyes," as the character is called by breeders, is quite common 

 in English Herefords, about two-thirds of the cattle showing it (out of 

 110, 73 showed more or less colour about their eyes), though it has 



