12 



About the time the mitochondrial mass begins to slip in 

 between the nucleus and the capsule, one or two deeply stain- 

 ing granules appear on the border line between the nucleus 

 and the capsule (Figs. 44 to 48). Koltzoff '06 in his re- 

 searches on the spermatogenesis in Galathea squamifera has 

 identified these granules with the centrosome. In my prepara- 

 tions of Menippe meroenaria I am able to distinguish the centro- 

 some for some time after the second maturation division (Pigs. 

 28 to 3E), but in the later resting period of the spermatid 

 and in the stages during the origin of the capsule, I am unable 

 to distinguish any granule that can with any certainty be iden- 

 tified with the centrosome » I shall call the structure develop- 

 ed from this granule, the central body . I am unable to follow 

 the development of two distinct granules although two could 

 some times be clearly distinguished as shown in ?ig. 44. Prob- 

 ably only the outer one is concerned in the development which 

 is here presented. 



This outer granule elongates (Fig. 47) and becones tubular 

 (Figs. 50 to 56). There soon appears at the outer end a vesicle 

 which Increases in size as the central body elongates. yiThile 

 the vesicle is still small there appears in its outer wall a 

 flattened granule which Is usually seen to be connected with 

 the end of the central body by means of a fine strand as though 

 it might have been derived from this body. As the central 

 body Increases in length and the vesicle enlarges its outer 

 wall approximates the outer wall of the capsule. The deeply 

 staining substance in the outer wall of the vesicle now becomes 

 connected with the wall of the capsule. (Pig. 56 to 58). A sec- 



