LIXE^E 287 



LINE/E. 



Benth. et Hook. Gen. PI. L 241. 



Fruit and Seed. The ovary is superior, syncarpous. and 

 typically five-celled, but often becoming spuriously ten-celled 

 by an outgrowth from the inner face of each carpel. Excep- 

 tions to the general rule occur in Eadiola millegrana, where the 

 flower is tetramerous. The ovary of Eeinwardtia tetragyna is 

 also typically four-celled, but that of E. trigyna and Anisadenia 

 is three-celled. The ovules are geminate and collateral, rarely 

 superposed, suspended from the inner angle of each cell near 

 the top, and are anatropous, with the micropyle superior, 

 and the raphe ventral. The loculi of several species of 

 Erythroxylon and Durandea are one-ovuled. 



The fruit is a capsule breaking up into as many cocci as 

 there are carpels, and dehiscing along the ventral suture or 

 separating along the dorsal suture, thus forming twice as many 

 pieces as there are carpels. It is membranous, one-seeded by 

 abortion, and said to be indehiscent in Anisadenia. The fruit 

 of the tribes Hugoniese and Erythroxyleae is drupaceous or 

 baccate, with as many pyrenes as there are carpels, or with 

 one only by suppression and then one-seeded. The fruits of 

 the Ixonanthese are subdrupaceous, but ultimately dehisce at 

 the apex. The cocci of Phyllocosmus are not septate. The 

 seeds are solitary or geminate in each cell, pendulous, anatro- 

 pous, often compressed laterally with the ventral edge and raphe 

 to the placenta, obovoid or oblong, with the testa sometimes, 

 though rarely, drawn out into a membranous wing. The 

 endosperm is fleshy or cartilaginous, copious, scanty, or alto- 

 gether wanting ; and the embryo is straight or rarely incurved, 

 almost equalling the cavity in length ; the cotyledons are flat, 

 plano-convex, ovate, elliptic, or linear, and foliaceous, and the 

 superior radicle nearly equals them in length, with some 

 exceptions such as Anisadenia and Sarcotheca, in which it is 

 short. 



The seeds are always laterally compressed when the car- 

 pels number three, four, or five with twice as many seeds, but 

 when the carpels are reduced to one, they are thick, obovoid, 



