PROTEACEJE 449 



The Order is divided into two series according as the fruit 

 is nut- or drupe-like and indehiscent ; or dehiscent and capsular 

 or follicular ; but in the latter group there are some exceptions. 

 The dry woody fruits may hang on the plants for years before 

 bursting open. 



The fruit of Grevillea Hilliana is a woody, one- to two-seeded 

 follicle, dehiscing by the separation of the two carpels. The 

 seed (fig. 628) is oblate, or oval, laterally much compressed, 

 winged all round and attached to the placenta by one edge. 

 The chalaza is at the upper end of the portion containing 

 the embryo, and the raphe pursues a circuitous course over 

 the wing, and then recurves towards the hilurn below the 

 middle of one edge of the seed. The embryo occupies the 

 whole of the interior, and has broadly obovate cotyledons 

 tapering to the base where they are strongly auricled in order 

 to accommodate the short radicle. 



The fruit of Hakea acicularis is broadly ovoid, thick- walled 

 and very woody, but dehisces by two valves when mature 

 and dry. When two seeds are produced in a fruit, they are 

 semi-obovoid by mutual pressure, gibbous on one side, and 

 flat or slightly concave on the faces in contact. The testa 

 is tuberculated and black, and is produced into a brown or 

 black, membranous and perforated wing, along one side and 

 round the apex of the seed. The wing is obliquely oblong- 

 ovate, about 1-5 cm. long, and 1 cm. wide. The cotyledons 

 are obovate, deeply auricled at the base, rather unequal in 

 thickness owing to mutual pressure and the peculiar form of 

 the seed. 



Seedlings. There is great variation among the cotyledons, 

 due chiefly to the size and shape of the seed in which they are 

 formed, and that again owes its shape to the cavity of the 

 fruit and whether one or two seeds are present. A large 

 number of them have cotyledons varying between roundly 

 obovate and oblong-obovate, auricled at the base, sessile or 

 subsessile, indented on one side or having a sinus which gives 

 them a falcate appearance. The roundly obovate form is well 

 represented by Hakea acicularis (fig. 631). The presence of 

 the auricles is explained in the account of the seeds of Grevillea 

 Hilliana. The leaves of Hakea acicularis are narrowly linear 



II. G G 



