JUGLANDE^E 519 



as long as the nucellus, and wide open at the apex. The colourless 

 tissue on each side at the base of the ovule has not much altered, 

 but vascular tissue begins to make its appearance. 



About the first week in July there seems no difference in the 

 size of the ovule, but the fruit as a whole continues to grow rapidly. 

 The bracteoles originally near the base of the fruit have been carried 

 a long way up. The neck has also been shortened by the thickening 

 of its basal portion. The vascular tissue has increased. Other and 

 more important changes have also taken place. On each side of the 

 ovule two oblong curved patches of tissue are seen lying about half 

 way between it and the exterior of the fruit ; these are losing colour 

 in the same manner as the above-mentioned masses at the base of 

 the ovule. 



In transverse section colourless tissue, evidently becoming corti- 

 cal, is seen to surround the ovule. On each side of the ovule towards 

 the circumference of the fruit four irregularly circular pieces of 

 colourless or cortical tissue are seen. The ovule is oval in outline 

 in the transverse section. The framework of the fruit, which ulti- 

 mately becomes sclerenchymatous, already commences to show itself. 



The ovule by the 23rd of July has increased but little in size, 

 but differs somewhat in shape from that of earlier stages. The 

 principal alterations are in the cortical tissues. In longitudinal 

 section a large oval or elliptic mass of cortex is seen on each side of 

 the fruit, mostly above the level of the ovule and close to the rind. 

 Two large masses also extend from the ovule downwards towards 

 the base of the fruit. Bounding all these masses are bands of tissue 

 that ultimately become sclerenchymatous. The cortex is as yet 

 solid. The cavity of the ovary is small. The bracteoles forming 

 the wings have grown nearly to the top of the fruit on the 

 posterior side, while they descend towards the base on the anterior 

 side. 



In the transverse section at this stage four masses of cortex are 

 seen, two on the right and two on the left. 



A longitudinal section of the fruit about the 6th of August 

 shows two longitudinal masses of cortex and a transverse piece, 

 lying as it were across the top of the two. All are separated by a 

 framework of vascular and sclerenchymatous tissue, or tissue 

 rapidly becoming sclerenchymatous. 



In the transverse section in a much more advanced state taken 

 at the very base of the ovary, a central line of small solid masses of 

 cortex is seen. Eight and left the usual four masses of cortex have 

 now become almost completely absorbed, leaving strands and broken 

 pieces running across the cavity. 



