100 



Investigations m> the. Life-History 



TABLE VII. 

 FATS OF PYLORI c APPENDAGES. 



May to August, - 



Oct. and Nov., 



These figures indicate : 



1st, That in the early part of the season there is a marked difference 

 as regards the amount of intestinal fat in fish at the mouth and in fish 

 in the upper reaches. 



2nd, That in October and November the amount of fat on the append- 

 ages of the fish coming to the mouth of the river has greatly diminished. 



3rd, That the "winter salmon" appearing at this time, 72 and 79, i.e., 

 fish not ready to spawn till the next season, have an enormous accumu- 

 lation of fat on the intestine. 



Taking the averages from May to August of estuary fish and fish iu 

 the upper waters, it is found that 28-5 grm. of fat per standard fish is 

 used up. 



From these results it would appear that this intestinal fat is the first to 

 be drawn upon, and that it is used up more rapidly than the muscle fat. 



(b) Liver Fat. 



In many fish, such as the cod, the great accumulation of fat occurs iu 

 the liver. I have shown elsewhere ("Journal of Physiol. ," vol. xix, p. 1 72) 

 that as much as 67 per cent, of fat is found in the liver of the Gadidae. 



In the female salmon there is never such an accumulation of liver 

 fat, In 1895 the amountof fat in theliver of several fish was determined, 

 and no marked change was observed from July to October in fish in the 

 estuary. 



In 1896 the liver fats were determined in four average fish : 



TABLE VIII. 



