104 Investigations on the Life- History 



The fat balance in male fish is given below : 



A. Balance, to August. 



Muscle. 



Gnus. 



Average in estuary fish from May to August, . .756 

 Average in upper-water fish in July and August, . 428 



Loss, . 328 

 Testes. 



Grms. 



Average iu estuary fish from May to August, . . 0'74 



Average in upper- water fish in July and August, . 2*41 



Gain, . 1'67 



The testes thus accumulate 1'67 grins, of fat, and thus 326 grms. of 

 fat are available as a source of energy. 



B. Balance to November. 



Muscle. 



Grms. 



Average in estuary fish in May arid August, . . 756 

 Average in upper- water fish in October and November, 103 



Loss, . 819 

 Testes. 



Grms. 



Average in estuary fish in May and August, . . 0*74 

 Average in upper- water fish in October and November, 6- 34 



Gain, . 5-46 



The testes take up 5-46 grms. of fat, and thus 813-5 grnis. are left 

 as a source of energy. 



On comparing these figures with the results obtained from female 

 salmon it will be seen that the accumulation of fat in the muscles is as 

 great in the male as in the female, and that the fat is used up to quite as 

 great an extent. On the other hand, the accumulation of fat in the testes 

 is trifling in amount, and thus the conclusion is indicated that in the 

 male the utilisation of fat as a source of energy is much greater than in 

 the female. This is especially marked in the later months. 



INTERNAL FAT. 

 A. Pyloric Appendages. 



The storage of fat on the pyloric appendages of male fish was not 

 inve>tigated by analyses. 



Some observations were, however, made upon the fats of the liver 

 during the season 1895. 



Table XIV. gives the results of these :- 



[TABLK. 



