1914] Sharp: Diplodinium ecaudatum 107 



maBig einzelne starkere quere Fibrillen (Fig. 37a)." Evidently there 

 is some mistake here for in "Fig. 37" the "a" refers to the inner lip 

 and apparently the walls in this region are structurally the same as the 

 walls in practically all other parts of the body. And in speaking of the 

 double fastenings of the dorsal membranelles (p. 158) he notes, that, 

 after penetrating the outer layer, their inner ends are connected with 

 the " Fibrillenschicht " by short inner supports, while to the outer 

 boundary layer decidedly longer "Fibrillen" proceed (Figs. 36, 37, 

 In. st.: aus. st.). And to quote from his description of the outer adoral 

 lip (pp. 158-159), "Die sie umgebenden Wiilste zeigen noch starkere 

 Fibrilleneinlagerung wie die des queren Membranellenzugs. ' ' He says 

 further, the insertions of adoral membranelles are similar to those of 

 the dorsal membranelles, viz., the double fastening, to the "Fibrillen- 

 schicht" internally and to the "Grenzschicht" externally. Then fol- 

 lows a detailed description of a very complicated fibrillar system 

 which may be rather briefly summed up as follows : These Stiitzfibril- 

 len," as Braune terms the ciliary roots, which here lie in the so-called 

 "Entoplasma a," are extraordinarily lengthened out. The innermost 

 set of these parallel "Fibrillen" (Fig. 39, St. fbr.) extend anteriorly 

 beyond the " Stiitzapparat " and in consequence of its spine-like pro- 

 longations are brought together in a circle. Near to the mouth opening 

 these fibrillae are in turn encircled by a " Fibrillenschlundring (Fig. 

 39, Schlr.)." On their oral ends are imposed the oral cilia and there- 

 fore they may be counted as "inneren Stiitzen" of the oral mem- 

 branelles. The outer supports (AuBenstiitzen) of these membranelles 

 likewise unite by threes or fours into " Stiitzfibrillen " which again 

 combine to form larger groups. Each of these larger groups has only 

 one point of attachment, which is found either in the "Stiitzapparat" 

 itself or upon the two " Fibrillenstammen " situated just above the 

 dorsal membranelle zone (Fig. 39, v auf &). 



This double fastening of the membranelles, viz., anterior and pos- 

 terior roots in the case of the dorsal membranelles and internal and 

 external roots in the case of the adoral membranelles, has been de- 

 scribed for D. ecaudatum in the main body of my paper, but these 

 fastenings were regarded by me as ciliary root-filaments rather than 

 as supporting fibrillae. An examination of figures 33 and 23 to 29 

 will show that in D. ecaudatum the only fibrillae present in the adoral 

 region are those which are imbedded in the oesophageal walls, and 

 according to Braune 's description such a picture as is obtained in the 

 contracted forms (text fig. D) would be absolutely impossible. 



