324 Forms of Reduplication 



It is also noteworthy, that complex reduplications may arise owing 

 to the combination of a primary and a secondary reduplication in the 

 same gametic series; e.g. the primary reduplications may be of the 

 following types : 



A and B = ?? : 1 : 1 : n 



A and C =m -.1 :1 : m, 

 B and C = p : \ : \ : p. 



The first two alone involve a secondary reduplication between B and C 



of the type 



nm + 1 : n + m : n + m : nm + 1, 



and this combined with the primary reduplication between B and C 

 gives the complex reduplication for B and C of 



BC : Be : bC : bc :: ^ {nm + 1) : n + ni : n + m : p (nm + 1). 



A careful study of systems of segregation will therefore, as soon as 

 two or more reduplications have been discovered, furnish the student of 

 genetics with data which will enable him on the one hand to test his 

 hypotheses by further experiment, and on the other hand to extend and 

 facilitate his researches. 



It must be borne in mind however that cell-divisions, if they do 

 really set up the phenomena of reduplication, must themselves depend 

 upon the structure of the protoplasm. It may be that the systems of 

 segi'egation will prove of some value in the analysis of this structure. 



University College of South Wales 

 AND Monmouthshire, Cardiff. 



