EANUNGULAGE^. 51 



adnate cells, dehiscing longitudinally by nearly marginal clefts.' The 

 carpels, also very numerous, are each composed of a unilocular ovary, 

 surmounted by a style grooved vertically along the whole of its inner 

 border, and slightly dilated at the tip. The whole of the upper part 

 of its lips is covered with stigmatic papillae. In the inner angle 

 of the ovary is a placenta which bears a fertile descending ovule 

 with its micropyle upwards and inwards ; and above it, in two ver- 

 tical rows, are a few- sterile ovules reduced to minute cellular nuclei. 

 The fruit is multiple, consisting of as many achenes as there were 

 carpels ; the fleshy albumen-^ of the seed surrounds a minute embryo. 

 In other species of this genus, such as Traveller's Joy ( C. Vitalba 

 L., Fr. Herhe mix Gucux) the 

 flower may equally consist of 

 four sepals, or it may have 

 five, six, or more. From six 

 to eight or ten are almost 

 constantly found in the beau- 

 tiful large-flowered species 

 cultivated in our conserva- 

 tories, as C. Jcuinginosa, patens, 

 fiorida, &c. The aestivation 

 is on the whole the same as 

 in C. Vitalba, but the thin 



inflexed portion of the sepal is here much broader.' We find 

 the same condition in C. Viticella L., and the other species which 

 have been united with it into a special section.'^ They are also dis- 

 tinguished by another feature ; the achene is only surmounted by a 

 short point (fig. 92) formed by the persistent base of the style. In 

 the other species, such as C. Vitaltja, the style persists on the summit 



' In C. Viticella the lines of dehiscence 

 are slightly internal; so too in C. Vitalba. 

 They are decidedly lateral in C. cirrhosa ; iu- 

 trorse in Atragene and Naravelia. Messrs. 

 Bentham & Hooker say (GeM. i.): " Antherce 

 introrsum dekiscant in Clematidihus 2 in- 

 dicis." 



2 The number varies; there are usually four 

 in two vertical rows ; more rarely two only, or 

 si.\ or eight. The superior ovules are always the 

 least developed. M. Rcepek saw, in 184'J (Bot. 

 Zeit., 1852, col. 187), four ovules in C.infegrifolia. 

 Payee was the first to show, in C. calycina, the 

 order of the evolution of the five ovules {Organog., 



253, t. 58). The existence of these abortive ovules 

 is another mark of the resemblance between Cle- 

 matis and Anemone. 



^ Its consistency varies; it may even become 

 quite horny. 



■• It is especially in these that we have imbri- 

 cation after the opening of the flower. (See note 1, 

 p. 50.) 



* Viticella DiLL., Nov. Gen. Giess. 165. — 



Spach, Suit, a Buff. vii. 



-Sect. 



DC, 



Sgsf. i. 160, Prodr. i. 8. Viticella, admitted as a 

 genus by Seki>'GE [Fl. des Jard. &c., iii. 80), 

 includes Clematis Viticella, Viorna florida, 

 cceridea and cylindrica. 



e2 



