ROSACEA. 



425 



Acioa ffuianensis. 



but the existence of a rudiment of this internal process of the 

 ovarian wall in Couepia proves the close relationship of that genus to 

 Parinari and the little value of the latter one.' 



In the Coupis^ (figs. 502, 503) we find the receptacle of llirtcHa 

 Couepia and most species of Parinari, with its deep unilateral tube. 

 In the flowers of Acioa ffuianeiisis, the pro- 

 totype of this genus, we find five imbricated 

 sepals, and as many alternating imbricated 

 petals. There are from ten to fifteen fer- 

 tile stamens all on one side of the flower; 

 on the other, that of the receptacular 

 tube, the androceal ring only bears a 

 variable number of little sterile teeth. 

 But the filaments of the fertile stamens 

 are united together for a long way up into 

 a long strap-like band, spirally involute 

 in the bud, and then spreading a long 

 way on one side of the expanded flower. 

 Towards the apex the filaments become 

 free and support each a short introrse two- 

 celled anther dehiscing longitudinally. 

 The gynseceum is inserted and formed as 

 in HirteUa. Its very long basilar style 

 ends in a scarcely dilated stigma ; it is 

 rolled up in the bud and exserted on an- 

 thesis. The fruit is a drupe with a usually dry hard thick mesocarp 

 and a one seeded endocarp ; the radicle of the embryo is inferior. 



Fig. 502. 

 Longitudinal section of flower (^). 



1 We have been unable to study the genus 

 Trichocarya (Miq.,Fl. Ind.- Bat., i.3o7; Suppl., 

 i. 116), which Bentham & Hooker place 

 {Gen., 607, n. 6) beside the genus Parinari, and 

 which, with the characters of the last species, a 

 pentameroua flower, an androceum of about 

 twenty-five stamens inserted all round the flower, 

 is said to present a tubular, cylindrical receptacle, 

 entirely filled up by an accrescent gynophore. The 

 ovary, inserted on top of the receptacular tube, 

 is described as containing one (?) ovule, and as 

 accompanied by a basilar style. The fruit is a 

 drupe, with a coriaceous endocarp lined with 

 hairs. Two species of this genus have been de- 

 scribed, one from Borneo, the other from Su- 

 matra, named by KORTHALS {Ned. Kruidk.Arch., 

 iii. 384, 388) Angeles>a splenden-s and Diemenia 



racemosa respectively. Were it not for the sin- 

 gular character of the gynophore, which, accord- 

 ing to MiQUEL himself, needs verification, it 

 would seem that these plants should be referred 

 to the genus Parinari. In his last work on these 

 plants MiQUEL {Ann. Mus. Lugd.-Bat., iii. 236) 

 has maintained as distinct genera Angelesia and 

 I>iemenia, which, as we have just seen, had been 

 united by Bentham & Hooker in 1865. It 

 appears that the herbarium at Leyden does not 

 contain enough specimens to allow any clear and 

 definitive judgment to be given on these points. 

 2 Acioa Atjbl., Otiian., 698, t. 280.— DC, 

 Prodr., ii. 526.— Spach, Suit, a Biiffon, i. 371. 

 — H. ')&lS.,Adamonia,\\. •221.— Ada (part.) W., 

 loc. fit. — Duhicia Neck., Elem., n. 1236 (ex 

 DC.).— Couepia (part.) B. H., Gen., 608, n. 9. 



