138 MISCELLANEOUS STUDIES 



Subfamily Pemphiginae Mordwilko 25 



Mordwilko, Ann. Mus. Zool. Imp. Acad. Sci. St. Petersburg, vol. 13, pp. 

 362-364, 1908. 



A summary of Mordwilko 's description of this subfamily has 

 already been given. The latest and probably the most complete sys- 

 tematic work on this subfamily that has been done is that of Dr. Albert 

 Tullgren of Stockholm, Sweden, in his paper, ' ' Aphidologische 

 Studien I" in 1909. Tullgren divides this subfamily into six tribes, 

 viz: Vacunina, Hormaphidina, Mindarina, Pemphigina, Schizoneu- 

 rina, and Anoeciina. In the tribe Vacunina he places Vacuna Heyden 

 and Glyphina Koch ; in Hormaphidina is the one genus Hamamelistes 

 Shimmer; in Mindarina is the one genus Mindarus Koch; in Pem- 

 phigina he places Asiphum Koch, Pachypappa Koch, ProciphUus 

 Koch, Thecdbius Koch, and Pemphigus Hartig; in Schizoneurina he 

 places the two genera, Schizoneura Hartig, and Tetraneura Hartig; 

 and finally in the Anoeciina is found the one genus Anoecia Koch. It 

 can be seen that he uses several of Koch 's genera which have not here- 

 tofore been generally used, namely: Prociphilus Koch, Thecabius 

 Koch, Asiphum Koch, and so forth. Lately there has been a tendency 

 among American aphidologists to accept these genera, and thus to 

 divide up the larger genus Pemphigus into these smaller ones. Mord- 

 wilko in his keys divides this subfamily into four groups, namely: 

 Hormaphidina, Pemphigina, Schizoneurina, and Vacunina. In Hor- 

 maphidina he includes besides the genus Hamamilestes Shimmer, the 

 genera Hormaphis Osten-Sacken and Cerataphis Lichtenstein. In 

 Pemphigina he includes Pentaphis Heyden, Tetraneura Hartig, 

 Pemphigus Hartig, Aploneura Passerini, Rhizoctonus Horvath, and 

 Paracletus Heyden. In Schizoneurina he places Lowia Lichtenstein, 

 Colopha Monell, Pachypappa Koch, Schizoneura Hartig, Anoecia, Koch, 

 and Mindarus Koch. In Vacunina he includes but the one genus 

 Vacuna Heyden, which he does not separate from Glyphina Koch. 



There is considerable difference in the classifications of these two 

 authors, but as far as we are concerned here in California our genera 

 are placed about the same by both. Following is a translation of 

 Mordwilko 's key to the groups : 



2", The author has under way a more exhaustive study of this subfamily, par- 

 ticularly of the species of Pemphigus and Procipliilus. As this research is still 

 in progress, however, it was thought best to omit any report of it, the author here 

 confining himself merely to the records of the presence of the various species in 

 California. It was hoped to have this study completed at the present time, but 

 the unprecedented conditions of this season have made it necessary to delay further 

 study for the time being. 



