THE CEREBELLAR CORTEX 



39 



THE GRANULAR LAYER OF THE CEREBELLUM 



The second, or granular, layer of the cerebellar cortex has about the same thickness as 

 the molecular layer; viz. i millimetre. It is made up of very small cells, 5 to 10 /* in 

 diameter (G), which are polygonal and have a number of very short dendrites. These cells 

 are very difficult to stain. 



Plate XIX. shows three such cells of various shapes and their short, crenated dendrites, 

 which terminate near to the cell in a club-shaped extremity. Kolliker has seen these ends 

 terminating in a fine brush of fibres. Cajal has shown that these cells possess a neuraxon, 



FIG. 6. 

 FIG. 6. Diagrammatic representation of a section through the cerebellar cortex. The cells are reproduced from the plates. 



/, molecular layer. //, granular layer. ///, white matter. P, Purkinje cell with its neuraxon, /, entering the white matter. S, small 

 stellate cells of molecular layer. B, large stellate cells with basket fibres, b. These basket fibres surround the body of the Purkinje cell shown 

 in dotted outline. G, cells of the granular layer with long, straight neuraxon, g, ascending to the molecular layer, and there bifurcating to become 

 tangential fibres. These fibres run at right angles to the plane of section of the plate. M, moss-like termination of white fibres, m, entering the 

 cerebellum from without. //, large Golgi cell of the second type with dendrites in both granular and molecular layers and neuraxon dividing and 

 subdividing within the granular layer. /, terminal filaments and fibres, ', entering the cerebellum from without and ending around the branches of 

 the Purkinje cells. 



which is directed toward the molecular layer (g), and after entering it divides in a T-shaped 

 pair of branches, which pass as horizontal and longitudinal fibres in this layer. In the plate 

 one such neuraxon is seen ascending from the middle cell toward the molecular layer. This 



