92 Experiments with Primula sinensis 



individual. The effect of the large yellow eye in giving rise, in the 

 absence of the factor for short-style, to the " homostyled " form has 

 been fully described on previous occasions'. Both the large yellow eye 

 and the white eye have effects when combined with certain colour 

 characters of the flower. Certain coloured forms possess a blotch of 

 deep colour, which in flowers with the ordinary eye occupies a well- 

 defined area at the base of the corolla lobes (Plate XXXI, figs. 50, 51). 

 If this character be combined with the large yellow eye, the deep colour 

 is, so to speak, pushed further outwards, and forms a rather ill-defined 

 band round the periphery of the area occupied by the pigment of the 

 eye^ But, so far as my observations go, when "Queen Alexandra" is 

 crossed with the same coloured race, the blotch of deep colour is 

 not developed in the F^ plants which have the white eye, though the 

 corresponding forms with the ordinary eye are blotched. 



(1) Large yellow eye x small eye. 



The accompanying table (p. 93) shows the results, inclusive of 

 those previously published', which have been obtained from crosses of 

 the " homostyled " plants with both short- and long-styled plants having 

 the ordinary eye. The crosses in which the F^ plant was selfed show 

 a considerable deficiency of large-eyed offspring, and in those cases in 

 which the small-eyed parent was short-styled, the deficiency is almost 

 confined to the short-styled offspring. The crosses of the form 

 {DR X R) have given results which, in the aggregate, do not differ 

 appreciably from expectation, though again, in those cases where we 

 are also concerned with short and long st)'le, the distribution of the 

 offspring among the four types is not very smooth*, and is particularly 

 irregular in one aberrant family (given separately in the table) where 

 the excess of short-styled offspring with the small eye is very marked. 



It is only in the early years, however, that any great discrepancy 

 manifests itself Very few crosses have been made with short-styled 

 parents since 1906 ; but experiments with long-styled plants have been 



^ Bateson and Gregory, loc. cit. pp. 582 — 584. 



* For illustration of flowers of this kind see Bateson, MendeV» Principles of Heredity, 

 Camb. Univ. Press, 1910, Plate VI. figs. 19, 21. 



^ Bateson and Gregory, he. cit. p. 584. 



•* In this connexion it must be borne in mind that in the crosses between short and 

 long style there is throughout a deficiency of short-styled offspring when the f j is selfed, 

 and an excess when the Fj is crossed with the long-styled. This would, of course, 

 have a disturbing efifect in cases such as that under notice. 



