114 Experiments with Primula sinensis 



are dark reds, of which a few approximate to "Crimson King," terra- 

 cottas of two shades, one bluer (Plate XXXI, fig. 47), the other a clean 

 red (fig. 48) and light reds corresponding with both the shades of 

 terra-cotta ; in the magenta class a similar series of forms occurs. The 

 grading between the sub-classes is close and I am not able to give any 

 precise numerical results as to the proportions of the various types. 

 The distribution of the green and red stigma among the magentas and 

 reds clearly indicates the existence of partial gametic coupling between 

 the factors for magenta and green stigma (see p. 127). 



^3 families have been raised from certain of the F^ forms in the 

 hope of elucidating their relations to one another and to the " Straw- 

 berries." The bluer terra-cotta appears to be differentiated from the 

 red kind by the addition of a single factor, but for the most part the 

 results are complex and further data are required for their detailed 

 analysis. One result, however, is of interest in connexion with the 

 relation between the magenta and red colour. An F^ plant with 

 peculiar deep rosy flowers and red stigma, when selfed, gave forms 

 like itself and strawberries ; a light red with green stigma, self- 

 fertilized, gave light reds, terra-cottas of both shades, and strawberries, 

 all with green stigma. The two plants were crossed together recipro- 

 cally, and the two families thus obtained consisted of typical magentas, 

 reds (including light reds and terra-cottas) and strawberries, all with 

 green stigma. 



"Orange King." (Plate XXX, fig. 8.) "Orange King" originated 

 with Messrs Sutton as a sport from a strain of " Crimson King " ; it bred 

 true from its first appearance. The F^ from the cross with " Crimson 

 King" bears an exceedingly close resemblance to the latter; the mature 

 flowers of the hybrid are probably not to be distinguished from those 

 of the pure race, but in the young flowers there is a slightly more 

 magenta tint than in the pure strain of " Crimson King " with which I 

 have worked. In the F^ from this cross there were obtained 55 plants 

 like the F-^, and 14 "Orange King"; some very slight differences in 

 the depth of the colour were noticeable among the latter. The ex- 

 tracted "Orange King" had the true red stem-colour, as compared 

 with the purplish-red colour of the forms resembling " Crimson King." 



" Orange King " x " Snowdrift." The F^ of this cross is indistin- 

 guishable to the eye from that of the crosses of either the Rosy- 

 magenta or "Crimson King" with "Snowdrift." The constitution of 

 the F^ follows the general lines of the F^ from ("Crimson King"x 

 "Snowdrift") but is of course rather more complex, since the ^i is 



i 



