72 Zoological Society. 
face almost flat in the female Troglodytes Savagei, and it is even con- 
cave in the great males; whilst in both adult males and females of 
the Troglodytes niger it is convex. 
There are specific distinctions in the interior of the cranium of 
the two species: the olfactory (rhinencephalic) fossa closed by the 
cribriform plate, though very little wider, is considerably deeperin 
Troglodytes Savagei than in Troglodytes niger ; and the ‘ crista galli,’ 
which is small in Troglodytes niger, is absent in Troglodytes Savagei, ou 
nor is there any ridge continued from the fossa upon the inner sur- om 
face of the frontal in the line of the frontal suture. 
In Troglodytes niger there is a short ala minor sphenoidei continued ai 
outwards from the anterior clinoid process, and the upper and outer 
angle of the foramen lacerum anterius is produced into a short cleft : 
in Troglodytes Savagei the rudiment of the ala minor terminates at — 
the upper border of the foramen lacerum anterius, which has a sub- | 
quadrate form, and is not extended outwards into an angular fissure. 
The sella turcica is relatively shallower in Troglodytes Savaget than. 
in Troglodytes niger, in which it is shallower than in Man. 
Many other minor differences were noted, but these would be. rf 
better understood by the aid of the figures in the memoir. Some 
scepticism, the author observed, might be expected as to the“alleged 
specific distinction of the large and small chimpanzees by natural- 
ists who had not been able to realise the differences by actual 
comparison of the specimens; but Professor Owen felt no doubt 
that, as in the case of the Pithecus Morio, more extended knowledge 
of the new species would confirm the validity of its distinction from 
the Troglodytes niger. 
The stronger zygomatic arches and the more developed sagittal 
and lambdoidal crests might be viewed as adaptive developments 
concomitant on the larger canines, and indicative of a larger and 
more powerful variety of chimpanzee; but the larger proportional 
molars and the smaller proportional incisors, the more equal and — 
complex last molar tooth, together with the prominence—slight as 
it is—of the nasal bones at their median coalescence, their inferior 
expansion, and, above all, the reappearance of the premaxillaries by 
their expanded superior extremities upon the face above the nostril, 
are more than mere differences of size and proportion, and being 
repeated in both male and female adults of the great chimpanzee of 
Gaboon, leave no alternative, according to the value assigned to 
such characters in other Quadrumanous genera, than to pronounce 
se 
the Troglodytes Savagei to be specifically distinct from the Troglo- 
dytes niger, and this to be, as the Pithecus Morio is to the Pithecus 
Wurmbii in Borneo, a smaller, feebler and more anthropoid species 
of the genus Troglodytes in Africa. 
In conclusion, Prof. Owen remarked that he had proposed the 
name of the new species of Chimpanzee provisionally, for the con- 
venience of its description and comparison ; and that, should he be 
able to learn that its discoverer had given a name to it, he should 
adopt that name, of which Troglodytes Savagei would then be a 
synonym. 
a «4 
The development of this crest also renders the supraoccipital sur- 
