Mr. J. Miers on several genera hitherto placed in Solanacee. 163 
nearly as possible within the limits prescribed by Mr. Robert 
Brown in his ‘ Prod.’ (doc. cit.), viz. to those genera with a mo- 
nopetalous corolla, with a 5-, rarely 4-partite border, even in ex- 
ceptional cases nearly regular and equal, the borders of whose 
lobes are always valvate or induplicato-valvate in eestivation ; 
epipetalous stamens, alternate with and equal to the number of 
the lobes, the fifth being seldom shorter and still more rarely 
sterile, anthers always bursting by longitudinal slits or pores ; 
an ovarium most generally 2-celled, rarely 3- to 5-locular, with a 
simple style, a bilobed or clavate stigma often hollow; a fruit 
either capsular or baccate, and albuminous seeds with a terete 
embryo, straight, and more or less curved in a nearly annular 
form, or somewhat spiral, the radicle in.all cases pointing to the 
basal angle of the seed, and turned away to some short distance 
from the hilum, which is generally lateral and marginal, rarely 
almost basal. 
The Scrophulariacee I would also propose should be confined 
to those genera that possess a tubular corolla more or less curved 
and irregular, with a 4- or 5-partite border generally unequal 
and bilabiate, the lobes rarely equal, but in every case with a de- 
cidedly imbricate zstivation ; stamens 2 or 4, didynamous, seldom 
with a fifth, which is very rarely fertile, often only rudimentary : 
an ovarium, most generally bilocular ; a simple style, with a stigma 
more or less bilabiate or bilobed ; the fruit almost always cap- 
sular (in very few instances baccate), 2-locular, rarely more-celled, 
bursting in various ways, with central placentz adnate to the 
dissepiment, and an embryo enveloped in albumen but little 
curved, generally with the radicle pointing to a basal hilum*; in 
one solitary instance (Campylanthus) the embryo is however peri- 
spherically curved. In this very natural family, although the 
floral leaves are often alternate, the cauline leaves are most gene- 
rally opposite, which occurs only accidentally in Solanaceae, and 
the origin of the inflorescence is strictly axillary. Thus limited, 
they form a very distinct natural order. 
The intermediate group, which I now propose as a suborder, 
under the name of Atropinee, or as a new order, under that of 
Atropacee, will consist of genera having a tubular persistent 
ealyx, more or less deeply divided, a hypogynous tubular corolla, 
with the tube more or less plicated in bud, and with a border. 
generally divided into 5 lobes slightly unequal, but which are 
* According to Mr, Bentham’s authority, DeCand. ‘ Prod.’ x. p. 186, and 
a statement positively affirmed by most botanists, but one which, it appears 
to me, must be received with some modification ; for in the seemingly truth- 
ful analyses of the genera figured by Nees v. Esenbeck, ‘Gen. PI. Germ.,’ 
the radicle is shown as in Solanacee, not pointing directly to the hilum. See 
plates of Erinus, Veronica, Wulfenia, Odontites, Euphrasia, Bartsia, Pedi- 
cularis, and Alectorolophus (Rhinanthus). 
11* 
